Phortica (Ashima) idiasta Gong, Zhu & Chen, 2018

Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru, 2020, Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species, Zootaxa 4789 (1), pp. 1-54 : 32

30) Phortica (Ashima) idiasta Gong, Zhu & Chen, 2018

Phortica (Ashima) idiasta Gong, Zhu & Chen, 2018: 440 .

Diagnosis. Pregonite partly less sclerotized (ch.43-0), not serrated on ventrodistal portion (ch.50-0); ventrosubapical portion of apical knob more or less protruded, forming barbule (ch.46-0); posterior postgonite dorsad curved, claw-like process, and anterior one basally divided into 3 long, outward oriented processes (“Figs 24, 25” in Gong et al. 2018).

Supplementary description. Supracervical setae approximately 14. Dorsomedial, tentorial apodeme 1/2 as long as basal, parallel portion of dorsolateral, tentorial apodeme. Arista with arb = 2/0; longest, dorsal branch as long as longest seta on pedicel; apical, leaf-like expansion narrower than long, apically roundish and with minute projection. Cibarial, medial sensilla 6–7 per side; posterior sensilla 3–4 per side.

The antisymmetry is observed in the phallic organs: in A-type, the posterior postgonite is present on the right side, the anterior one on the left side, and left gonocoxite of hypandrium more stretched out (“Fig. 25” in Gong et al. 2018); but in B-type, vice versa.

Specimen examined. Thailand: 1♂ (B-type), Umphang , Tak, 30.v.2008, H. Bänziger leg. ( SEHU) .

Distribution. China (Yunnan), Thailand *.

Remarks. This species first branched off in the glabra subgroup ( Fig. 1 ). The remaining three species form a cluster (BP = 96%) with three synapomorphies ( Fig. 1 ): pregonite ( Fig. 9G ) nearly entirely sclerotized (ch.43-1), only angulated in ventrosubapical portion of apical knob (ch.46-1), and coarsely serrated on ventrodistal portion (ch.50-1).

Figures

FIGURE 1. The strict consensus tree resulting from the parsimony analysis I (PAUP* v4.0a166) of the data matrix of 40 spp. × 66 morphological characters (Appendix 1) for the genus Phortica (especially focusing on the subgenus Ashima). Synapomorphies (solid circle: nonhomoplastic; open circle: homoplastic) inferred from both ACCTRAN and DELTRAN character optimization are indicated on each internal branch along with support values (bootstrap frequency %).

FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E.