Tanyproctus (Tanyproctus) arher Bezděk, Sehnal & Král, 2013

Bezděk, Aleš, Sehnal, Richard & Král, David, 2013, Tanyproctus (Tanyproctus) arher (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), a new species from the Socotra Island, Yemen, Zootaxa 3737 (2), pp. 191-196 : 192-196

publication ID

https://doi.org/ 10.11646/zootaxa.3737.2.7

publication LSID

lsid:zoobank.org:pub:BE471AA1-71E6-4B94-BD10-D48352E0C967

DOI

https://doi.org/10.5281/zenodo.5698259

persistent identifier

https://treatment.plazi.org/id/03C787B9-FFC0-EC37-FF7D-D457FBEFFED9

treatment provided by

Plazi

scientific name

Tanyproctus (Tanyproctus) arher Bezděk, Sehnal & Král
status

sp. nov.

Tanyproctus (Tanyproctus) arher Bezděk, Sehnal & Král , new species

( Figs 1, 2 View FIGURES 1 – 4. 1 – 2 , 5, 6 View FIGURES 5 – 8. 5 – 6 , 9 View FIGURE 9 )

Type locality. Yemen, Socotra Island, Hala [= Haalla] area, Arher , 12°33.0′N, 54°27.6′E, 5 m a.s.l., freshwater spring in sand dune.

Type material. Yemen ( Socotra Isl.): Holotype ♂ (NMPC): “ Yemen, SOCOTRA Island / Hala area, ARHER / freshwater spring in sand dune / 9–10.+ 15-vi-2012 / 12°33.0′N, 54°27.6′E, 5 m // SOCOTRA expedition 2012 / J. Bezděk, J. Hájek, V. Hula, / P. Kment, I. Malenovský, / J. Niedobová & L. Purchart leg. [printed label]”. Paratype ♂ [pinned specimen, left middle leg preserved in pure alcohol] (ZFMK): same data as holotype.

Description of holotype (♂). Body length 8.5 mm. Body elongate; surface dark brown to black coloured, appendages and sutural elytral interval somewhat lighter; shiny; macrosetation pale ( Fig. 1 View FIGURES 1 – 4. 1 – 2 ).

Head. Labrum small, bilobed; lobes round, coarsely, irregularly punctate. Outline of clypeus almost trapezoidal, with distinctly upturned margin; clypeus remarkably depressed along margin and excavated centrally; anterior margin shallowly emarginate, anterior angles round, sides broadly arcuate ( Fig. 2 View FIGURES 1 – 4. 1 – 2 ). Genae narrow, round. Frontoclypeal suture feebly arcuate, considerably impressed. Eyes relatively large, distinctly exceeding genae externally in dorsal aspect ( Fig. 2 View FIGURES 1 – 4. 1 – 2 ); distance between eyes in ventral aspect extended remarkably diameter of eye. Punctation of clypeus coarse and dense, almost evenly distributed but impunctate centrally; punctures separated by approximately their diameter, each puncture bearing short, semierect macroseta. Vertex rather rugopunctate; punctures separated by less than their diameter to confluent, each puncture bearing very short, erect macroseta; macrosetation becoming shorter posteriad. Antennae with 10 antennomeres; antennomere 2 short, approximately as long as wide; antennomeres 3–5 elongate. Club pentamerous, straight, shorter than antennal shaft (antennomeres 1–5 combined). Antennomeres 1–5 with sparse, long macrosetae, club with sparse, short macrosetae. Terminal maxillar palpomeres elongate, rounded apically, absent from depression, approximately of same length as palpomeres 2 and 3 combined.

Pronotum moderately convex, transversal, broadest just anterior to middle except broad basal interruption all around border; anterior bead flat, narrow, distinctly widened medially. Punctures irregular; each puncture bearing long, erect macroseta; macrosetae becoming regularly shorter mediad. Lateral margin considerably coarsely crenate, with row of long macrosetae; basal margin with row of finely and irregularly distributed punctures bearing long, recumbent macrosetae in posterior angles and short macrosetae medially. Anterior angles prominent, projecting over anterior margin, acute-angled, with round apex; sides in approximately anterior half almost straight, divergent posteriad to very broadly obtuse posterior angles; posterior margin broadly round. Surface glabrous and finely, coarsely, slightly irregularly punctate, punctures separated by 2–4 their diameters, area along medial longitudinal line smooth.

Scutellum approximately as wide as long, triangulate, sides broadly arcuate, apex acute, surface with several punctures basally and laterally.

Elytra convex, slightly dilated posteriad, sutural angle rounded; striae excepting sutural stria missing or only very feebly indicated; distinctly microsculptured with disc feebly transversally wrinkled; punctation coarse, dense, almost regular; punctures separated by 1–2 their diameters. Sutural interval slightly convex; sutural stria with row of irregularly distributed punctures; lateral margin distinctly bordered with row of long, erect setae.

Macropterous.

Legs. Femora moderately shiny; metafemora with entire surface sparsely, almost regularly punctate; punctures bearing long, recumbent macrosetae. Protibia tridentate; basal tooth considerably weak; terminal calcar long, sharp, slightly curved externally, acute apically, inserted against emargination between basal and medial teeth. Mesotibia and metatibia slightly expanded apicad, with two setiferous transversal carinae. Mesotibial terminal calcars equal in length, flattened, acute apically. Metatibial terminal calcars equal in length, considerably flattened, acute apically. Protarsomeres 2–4 considerably dilated, protarsomere 2 with length-width ratio: 1:2. Mesotarsomeres 2–4 remarkably more slightly dilated that those of protibiae; protarsomeres and mesotarsomeres 1-4 with shortly and densely macrosetaceous pads ventrally; metatarsomeres covered with long sparse macrosetae ventrally. Claws bifid.

Ventral surface of thorax with dense, long, recumbent macrosetation.

Propygidium microsculptured, coarsely, sparsely, irregularly punctate; pygidium finely microsculptured; border complete; surface coarsely and irregularly punctate.

Ventrit es almost bare, remarkably coarsely and irregularly punctate.

Male genitalia ( Figs 5–6 View FIGURES 5 – 8. 5 – 6 ). Aedeagus symmetrical; parameres slender, relatively long, only slightly shorter than phallobase, distal part not dilated in dorsal aspect.

Female. Unknown.

Varability. The only known paratype slightly differs in size (total body length: 8.4 mm) and with lighter coloured elytra.

Differential diagnosis. Tanyproctus (T.) arher is morphologically very similar to the other endemic Socotran Tanyproctus species (Král et al. 2012). In the key to males of Socotran Tanyproctini (Král et al. 2012: 180), T. (T.) arher will key to the couplet with T. (T.) wraniki . For differentiation from this species see the complex of diagnostic characters in Table 1 View TABLE 1 .

Sehnal & Bezděk, 2012

Etymology. Derived from area of origin of the new species, the Arher freshwater spring, Socotra ( Yemen); noun in appositon.

Collecting circumstances. Both specimens were captured at light at about 6 p. m. (J. Hájek, personal communication).

Geographical distribution. Endemic species to the Socotra Island; both so far known specimens originates from the coastal area of Haalla (E Socotra), for details see Bezděk et al. (2012).

TABLE 1. Differential characters of Tanyproctus (T.) arher Bezděk, Sehnal & Král, new species and T. (T.) wraniki Král,

species / character shape of clypeus Tanyproctus arher trapezoidal, longer (Fig. 2) Tanyproctus wraniki trapezoidal, shorter (Fig. 4)
sculpture of clypeus size of eyes excavate and impunctate centrally (Fig. 2) relatively large, considerably prominent laterad in dorsal aspect (Fig. 2) depressed and punctate centrally (Fig. 4) relatively small, only weakly prominent externally in dorsal aspect (Fig. 4)
metafemora entire surface sparsely, almost regularly punctate punctation coarser, somewhat irregular, restricted approximately to marginal areas, impunctate centrally
shape of protarsomeres 2–4 shape of parameres considerably dilated, protarsomere 2 with length-width ratio: 1:2 as in Figs 5–6 dilated, protarsomere 2 with length–width ratio: 1.0:3.7 as in Figs 7–8
body size distribution 8.4–8.5 mm E Socotra (Haalla coastal area) 6.8 mm (holotype) C Socotra (Diksam plateau)

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Melolonthidae

Genus

Tanyproctus

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