Heteroneda Crotch

POORANI, J., 2023, An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini, Zootaxa 5332 (1), pp. 1-307 : 149

Genus Heteroneda Crotch

Heteroneda Crotch, 1871: 6 . Type species: Coccinella reticulata Fabricius, 1801 (by subsequent designation of Crotch 1874: 162) (= H. billardieri Crotch, 1871 ; Iablokoff-Khnzorian 1982: 538).

Diagnosis. Elytra moderately to strongly convex, glabrous. Anterior clypeal border straight between lateral projections. Antenna 11-segmented; slightly shorter than head capsule, with a weak three-segmented club; terminal antennomere longer than penultimate, rounded apically. Terminal maxillary palpomere securiform. Pronotal disc evenly convex transversely, with narrowly upturned or slightly thickened external borders. Prothoracic hypomeron without foveae near anterolateral corners; prosternal process with distinct carinae extending forward to the middle of the prosternum. Anterior margin of mesoventrite distinctly, deeply emarginate medially. Lateral margins of elytra with complete beads; epipleuron without foveae. Tibial spur formula 0-2-2. Abdominal postcoxal line not recurved, incomplete laterally, without an oblique associated line (From Ślipiński et al. 2020).

Distribution. Oriental and Australo-Pacific regions.

Affinities. Tomaszewska et al. (2021) included it in the Synonycha- group of genera and recovered it as a strongly supported genus in a clade comprising Calvia (part), Singhikalia , Phrynocaria (including part of Coelophora ), and Docimocaria . Their analyses suggest Heteroneda is closely related to Phrynocaria , “albeit with weak support”.

Included species. The type species, H. billardieri Crotch , is distributed in this region ( Ślipiński et al. 2020).