Diplommatina godawariensis Budha & Naggs, 2017

Budha, Prem B., Naggs, Fred & Backeljau, Thierry, 2017, The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species, European Journal of Taxonomy 337, pp. 1-30 : 11-12

Diplommatina godawariensis Budha & Naggs sp. nov.

urn:lsid:zoobank.org:act:2ED01F49-7E8A-4A53-A021-720968DC721A

Figs 4B , 9E

Diagnosis

Measurements (n = 5): SH 3.2–3.5 mm, SW 1.7–1.8 mm, Wh: 6.5–7.0; holotype: SH 3.3 mm, SW 1.8 mm, Wh 7.0. Dextral, penultimate and body whorl nearly equal in width. Constriction at the inner edge of PeP ( Fig. 9E ). Two parietalis: Prt1 short and strong, Prt2 not very distinct, nearly attached to the sutural wall ( Fig. 4B 3 ). Two vertical palatalis, Vpt1 strong and runs along the constriction, Vpt2 short and small, difficult to distinguish, just above the outer lip of the parietal wall ( Fig. 9E ). Two horizontal palatalis: both Hpt1 and Hpt2 are thin, situated just above the columella, posterior longer than anterior ( Fig. 9E ). Columellaris distinct ( Fig. 4B 3 ). Columellar tooth prominent. Protoconch smooth, Wh approx. 2.0, with small pits evenly distributed, no radial ribs ( Fig. 4B 1 ). Teleoconch with radial ribs; about 5–6 ribs/ 0.5 mm on the penultimate whorl; 5–7 ribs/ 0.5 mm on the body whorl towards the aperture ( Figs 4B 2 , B 4 ). No spiral striation. Umbilicus closed. Peristome with two lips; outer lip slightly reflected; inner lip thick and forming a thin callus on the penultimate whorl. Angular edge of the peristome rounded. Basal edge of the peristome outwardly directed.

Etymology

The species name refers to the type locality ‘Godawari’.

Type material

Holotype

NEPAL: Godawari , Lalitpur District, 1551 m, mixed forest, 27.589284 N, 85.388309 E, 1 Dec. 2006, P.B. Budha leg. ( CDZMTU032 ).

Paratypes

NEPAL: 5 shells from the type locality, same data as holotype ( CDZMTU 033); 2 shells, Godawari, 1551 m, 27.5969 N, 85.3890 E, 30 Apr. 2007, P.B. Budha leg. ( CDZMTU 033b).

Remarks

Diplommatina godawariensis sp. nov. is similar to D. diplocheilus Benson, 1857 from Khasi Hills, NE India “Teria Ghat, Khasi Hills” ( Hanley & Theobald 1876: pl. 140, figs 2–3), but D. godawariensis sp. nov. has a smaller columellar tooth and a narrower penultimate whorl. The aperture of D. godawariensis sp. nov. is rather similar to that of D. pachycheilus , but this latter species has more tumid apical whorls and its penultimate whorl is slightly narrower than that of D. godawariensis sp. nov.

Figures

Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.

Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov.

Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.

Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal.

Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two).