Hlavaciellus smetanai, Jałoszyński, Paweł, 2011

Jałoszyński, Paweł, 2011, Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 2763, pp. 1-33 : 9-11

Hlavaciellus smetanai sp. n.

( Figs. 11 , 8 , 22, 23 )

Type material. Holotype: MALAYSIA: ɗ, "BORNEO SABAH \ Tamis, Hwy Al \ 10 km NW Kinabalu \ Park Entr. 1100 m \ 24.V.87 A. Smetana" [white, printed], " HLAVACIELLUS \ smetanai m. \ det. P. JAŁOSZYŃSKI, '09 \ HOLOTYPUS " [red, printed] ( MHNG).

Diagnosis. Head in males with broad and deep transverse, in middle angulate, groove running from posterior margin of each eye towards middle and only slightly anteriorly, remaining parts of head non-modified.

Description. BL 1.90. Body of male ( Fig. 11 ) strongly convex, pigmentation light brown, vestiture yellowish.

Head ( Fig. 8 ) large, HL 0.33, HW 0.50; vertex convex, with highest point located in middle of anterior part, anterior margin broadly V-shaped and bordered anteriorly by pair of broad and deep, transverse impressions running from posterior margin of each eye towards middle and only slightly anteriorly and forming entire (i.e., not disrupted in middle) groove; frons nearly pentagonal, non-modified, slightly flattened, confluent with subtrapezoidal, convex clypeus; supraantennal tubercles moderately raised and not separated from frons; eyes large, strongly convex and coarsely faceted. Punctures and setae as in Fig. 8 . Antennae as long as about 0.7 BL, slender, gradually thickened up to antennomere X, apical margin of antennomere IX and X slant (i.e., external lateral margin longer than internal one); antennae covered with thin, long, dense and suberect setae, with several longer, straight and more erect setae on antennomeres VII–IX, AnL 1.35, antennomere I 1.3x as long as broad; II distinctly narrower and shorter than I, 1.2x as long as broad; III slightly larger than II, 1.3x as long as broad; IV slightly larger than III, 1.2x as long as broad; V slightly larger than IV, 1.4x as long as broad; VI slightly larger than V, 1.4x as long as broad; VII minimally larger than VI, 1.3x as long as broad; VIII slightly broader but not longer than VII, 1.2x as long as broad; IX slightly larger than VIII, 1.1x as long as broad; X distinctly broader and slightly longer than IX, about as long as broad; XI slightly narrower than X and slightly shorter than IX–X together.

Pronotum in dorsal view subtrapezoidal with strongly convex anterior margin, nearly equally broad from base up to anterior third, PL 0.58, PW 0.75; sides strongly rounded in anterior third, then nearly straight and parallel up to distinctly sharp but rounded hind angles; posterior margin deeply bi-emarginate; ante-basal pits very shallow but relatively well marked, each located slightly closer to posterior than to lateral margin of pronotum; lateral edges slightly swollen on entire length to form narrow but distinct lateral carinae. Pronotal disc glossy, in middle covered with very fine and shallow punctures separated by spaces 1.5– 2 x as wide as puncture diameters, punctures close to margins of pronotum are larger and denser, those near each hind angle of pronotum are slightly coarse. Setae long, moderately dense, suberect to erect.

Elytra oval, relatively short, broadest between anterior third and middle, EL 1.00, EW 0.88, EI 1.14; basal pit on each elytron small but distinct, nearly adjacent to scutellum; subhumeral lines sharply marked, as long as 0.40x EL, only slightly divergent, each developed as border between higher humeral region and lower adsutural area; apices of elytra separately rounded. Punctures on basal third of each elytron small but more distinct than those on pronotum, gradually reducing in diameter and depth toward lateral margins and apices; setae distinctly longer, thicker and more erect than those on pronotum. Hind wings fully developed.

Legs slender and long, tibiae straight or nearly straight.

Metaventrite with shallow postmesocoxal impressions barely reaching half length of ventrite.

Aedeagus ( Figs. 22, 23 ) stout and relatively short in relation to body length, AeL 0.30; median lobe gradually narrowed towards subtriangular, pointed apex; internal armature nearly symmetrical; parameres in lateral view relatively slender, with very short curved apical parts.

Female. Unknown.

Distribution. East Malaysia: Borneo, Sabah.

Etymology. The name is dedicated to the collector of the holotype, a renowned specialist on Staphylinidae, Dr. Aleš Smetana.

Remarks. The modifications of the head in H. smetanai are similar to those in H. adelphos and H. bifoveatus . However, the clearly separated and densely punctated vertex and the deep, entire impression between the frons and vertex are sufficient to identify this species. The impressions in the two mentioned congeners are much shallower and distinctly separated in middle (in. H. adelphos ) or separated in middle and not reaching the posterior margin of eye ( H. bifoveatus ). Moreover, the very broad, stout aedeagus of H. smetanai is clearly different from very slender aedeagi of H. adelphos and H. bifoveatus . Interestingly, a very similar aedeagus can be found in H. metrios described below. However, the latter species belongs to the group characterized by simple, non-modified heads of males.

Figures

FIGURES 10 – 12. Dorsal view of holotypes of Hlavaciellus, males. 10, H. decoratus sp. n. 11, H. smetanai sp. n. 12, H. adelphos sp. n. Scale bars: 1 mm.

FIGURES 4 – 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm.

FIGURES 13 – 23. Aedeagus in ventral (13, 15, 18, 20, 22) and lateral (14, 16, 19, 21, 23) views, and head of male in lateral view (17). 13, 14, Hlavaciellus peninsularis sp. n. 15 – 17, H. cirrus sp. n. 18, 19, H. angustifrons sp. n. 20, 21, H. decoratus sp. n. 22, 23, H. smetanai sp. n. Scale bars: 0.2 mm (13 – 16, 18 – 23), 0.1 mm (17).