Colocasiomyia zeylanica (Okada, 1986)

Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak & Toda, Masanori J., 2023, Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species, Zootaxa 5278 (2), pp. 201-238 : 222

publication ID	https://doi.org/ 10.11646/zootaxa.5278.2.1
publication LSID	lsid:zoobank.org:pub:697AC989-141D-4A89-8F02-5A4644E303EA
DOI	https://doi.org/10.5281/zenodo.7906009
persistent identifier	https://treatment.plazi.org/id/03C687D9-FFC3-FFF5-A5C6-FF28845DE016
treatment provided by	Plazi
scientific name	Colocasiomyia zeylanica
status

Treatment

1. Colocasiomyia zeylanica View in CoL species subgroup, new

Diagnosis. Male sternites V and VI fused to each other (ch.46-1; Fig. 10A–C View FIGURE 10 1 View FIGURE 1 ). Hypandrial phragma vertical plate-like (ch.68-1; Fig. 11-1A–C 3,4).

Shared characters. Antennal arista distally narrowing (ch.4-0; Fig. 3A–C View FIGURE 3 , “ Fig. 1A, B View FIGURE 1 ” in Okada 1986, “ Fig. 6 View FIGURE 6 ” in Grimaldi 1991). Eye with stout, dense, interfacetal setulae (ch.7-1; Fig. 2A–C View FIGURE 2 1 View FIGURE 1 ). Proclinate, orbital setae present (ch.8-0; Fig. 2A–C 2 View FIGURE 2 , “ Fig. 1A, B View FIGURE 1 ” in Okada 1986, “ Fig. 6 View FIGURE 6 ” in Grimaldi 1991). Inner, vertical setae longest on head, in line with ipsilateral orbitals ( Fig. 2A–C 2 View FIGURE 2 ). Outer, vertical setae present (ch.9-0; Fig. 4A–C View FIGURE 4 , “ Fig. 1A, B View FIGURE 1 ” in Okada 1986, “ Fig. 6 View FIGURE 6 ” in Grimaldi 1991). Facial carina shorter than pedicel + 1st flagellomere ( Fig. 2A– C 2 View FIGURE 2 ). Gena blackish brown. Upper, postocular setae down-curved, except uppermost ones (ch.10-0; Fig. 4A–C View FIGURE 4 ). Patch of supracervical setae present between dorsomedial and -lateral, tentorial apodemes (ch.13-0; Fig. 4A–C View FIGURE 4 ). Dorsomedial, tentorial apodeme reaching 1/4 to vertex; dorsolateral apodemes divergent, nearly straight or distally slightly curved outward (ch.14-0; Fig. 4A–C View FIGURE 4 ). Palpus apically with single, prominent seta (ch.17-1; Fig. 5A–C View FIGURE 5 1 View FIGURE 1 ). Clypeus not thicker than 1/2 length of cibarium ( Fig. 5A–C View FIGURE 5 1 View FIGURE 1 ). Cibarial, medial sensilla in parallel rows as wide as sensilla campaniformia; posterior sensillum 1 per side (ch.20-0, ch.21-2; Fig. 5A–C View FIGURE 5 2 View FIGURE 2 ). Thorax glossy, blackish brown ( Fig. 14A–C View FIGURE 14 ). Postpronotal lobe with 2 prominent setae subequal in length (ch.25-0; Fig. 6A–C View FIGURE 6 ). Anterior, dorsocentral setae situated posteriorly to transverse suture (ch.26-0; Fig. 6A–C View FIGURE 6 ). A pair of prescutellar, acrostichal setae present (ch.27-0; Fig. 6A–C View FIGURE 6 ). Acrostichal setulae in 6 rows (ch.28-1; Fig. 6A–C View FIGURE 6 ). Costal setae in middle row from 2nd costal section to proximal portion of 3rd section all apically blunt, heavy, peg-like; sparse setae in upper row small, weak, trichoid (ch.30-0, ch.31-0; Fig. 8A–C View FIGURE 8 1 View FIGURE 1 ). Patch covered with only minute pubescence absent on anterodorsal portion of fore tarsomere I (ch.33-0; Fig. 9A–C View FIGURE 9 1 View FIGURE 1 ). Hind tibia without apicoventral, stout setae (ch.40-0; Fig. 9A–C View FIGURE 9 3 View FIGURE 3 ). Claws shorter than tarsomere V (ch.43-0; Fig. 9A–C View FIGURE 9 ). Abdominal sternites grayish brown; I narrow, transverse plate more or less protruded or expanded posteromedially (ch.45-1; Fig. 10A–C View FIGURE 10 ). Epandrium largely pubescent on dorsal to lateral portion; ventral lobe not differentiated (ch.53-0, ch.56-0; Fig. 11-1A–C 1). Surstylus plate-like, fused to epandrium but not to cercus (ch.59-0, ch.60-1, ch.63-0; Fig. 11-1A–C 1,2). Membranous aedeagus absent (ch.70-2; Fig. 11-1A–C 3,4). Phallapodeme fused to phallal sheath (ch.75-1; Fig. 11-1A–C 4). Hypoproct nearly entirely pubescent (ch.80-0; Fig. 12A–C View FIGURE 12 1 View FIGURE 1 ). Epigynium dorsally neither constricted nor separated (ch.81-0; Fig. 12A–C View FIGURE 12 1 View FIGURE 1 ). Hypogynial valves with trichoid sensilla on both basal and distal portions ( Fig. 12A–C View FIGURE 12 ).

Included species. Colocasiomyia zeylanica *, C. nepalensis *, C. pinangae sp. nov., C. besaris sp. nov., and C. luciphila sp. nov. *Two known species were assigned to this species subgroup based on their original descriptions and illustrations ( Okada 1986; Grimaldi 1991); although the two diagnostic characters are not confirmed for them, they share the following homoplastic synapomorphies with the other members of this subgroup: eye with stout, interfacetal setulae (ch.7-1); and claws shorter than tarsomere V (ch.43-0; “ Fig. 1A, C View FIGURE 1 ” in Okada 1986).