Sosane Malmgren, 1866

Imajima, Minoru, Reuscher, Michael G. & Fiege, Dieter, 2013, Ampharetidae (Annelida: Polychaeta) from Japan. Part II: Genera with elevated and modified notopodia, Zootaxa 3647 (1), pp. 137-166 : 148-149

Sosane Malmgren, 1866

Type species: Sosane sulcata Malmgren, 1866

Synonyms:

Sosanopsis Hessle, 1917 Mugga Eliason, 1955

? Melinnata Hartman, 1965 Muggoides Hartman, 1965 Sosanella Hartman, 1965 Genus A sensu Uebelacker, 1984

Generic diagnosis (emended). Prostomium with middle lobe delimited by incision, without glandular ridges. Buccal tentacles usually smooth. Lower lip usually crenulated. Three to four pairs of cirriform branchiae; pairs 1–3 in segment II, 4th pair, if present, usually in segment IV. Notochaetae in segment II of same size as following notochaetae, or absent. Nine to 13 thoracic uncinigers. Notopodial cirri absent. Last, second-, third-, or fourth-tolast thoracic unciniger with elevated, enlarged notopodia, sometimes connected by mid-dorsal ridge. Notochaetae of modified notopodia with hirsute tips. One or two intermediate segments. No abdominal rudimentary notopodia present. Uncini with numerous teeth in 3 or more rows.

Remarks. The diagnosis has been emended to account for the first Sosane species with elevated and modified notopodia in the second-to-last and fourth-to-last thoracic uncinigers, respectively. Furthermore, the position of the branchiae, and the number of intermediate segments are included.

The synonymies of the genera listed above, suggested by Jirkov (2001, 2011), are accepted. The characteristic Sosane -like modification of a posterior thoracic unciniger in these genera is considered synapomorphic.

Sosanopsis Hessle, 1917 differs from Sosane Malmgren, 1866 only by the absence of chaetae in segment II (“paleae”). This character is not considered valid for the distinction of genera (Jirkov 1994a, 2001, 2008, 2011; Reuscher et al. 2009; Parapar et al. 2012).

The monotypic genus Sosanella Hartman, 1965 was described with 13 thoracic uncinigers. Our examination of the holotype revealed that only 12 thoracic uncinigers are present. The species Sosanella apalea Hartman, 1965 is herein considered a junior synonym of Sosane wireni , Sosanella a junior synonym of Sosane .

The holo- and paratype of Melinnata americana Hartman, 1965 are incomplete, missing the modified segment. The species, and the monotypic genus Melinnata Hartman, 1965 , are therefore indeterminable. The possible synonymy of the genus with Sosane is doubtful.