Glyphidrilus, Chanabun & Sutcharit & Tongkerd & Tan & Panha, 2012

Chanabun, Ratmanee, Sutcharit, Chirasak, Tongkerd, Piyoros, Tan, Shau-Hwai Aileen & Panha, Somsak, 2012, Three new species of semi-aquatic freshwater earthworms of the genus Glyphidrilus Horst, 1889 from Malaysia (Clitellata: Oligochaeta: Almidae), Zootaxa 3458, pp. 120-132 : 131

Key to species of Glyphidrilus from Malay Peninsula

1 Gizzard in 7 or ½7–½8 or 7–8........................................................................... 2

- Gizzard in 8 or 8–9.................................................................................... 4

2 Gizzard in 7; wings in 21–26; clitellum in 18–30; spermathecae in 12/13–6/17............................ G. jacobsoni

- Gizzard in ½7–½8 or 7–8; wings; clitellum or spermathecae different............................................ 3

3 Intestine from 15; wings in 23–½32; clitellum in 16–32; spermathecae in 13/14–17/18........................ G. weberi

- Intestine from 14; wings in 22, 23–28, ½29, 29; clitellum in 17, 18–31, 32; spermathecae in 14/15–17/18..................................................................................................... G. peninsularis sp. n.

4 Gizzard in 8.......................................................................................... 5

- Gizzard in 8–9....................................................................................... 11

5 Genital markings absent; wings in 19, 20–25.................................................... G. quadrangulus

- Genital markings different.............................................................................. 6

6 Spermathecae absent; wings in 24, 25–31, 32; clitellum in 19, 20–35, 36, 37......................... G. vangviengensis

- Spermathecae present; wings or clitellum different........................................................... 7

7 First hearts in 7; wings in 18, ½18, 19–24, ½24; clitellum in ½13–34; spermathecae in 14–18............... G. kukenthali

- First hearts in 8 or 9; wings; clitellum or spermathecae different................................................ 8

8 First hearts in 8; wings in ½23, 23–27, 28; clitellum in 17, 18, ½18, 19–28, 29, 30, ½31; spermathecae in 14–17.... G. horsti

- First hearts in 9; wings; clitellum or spermathecae different.................................................... 9

9 Wings in 21–25, ½26, 26; clitellum in 18, 19–27, ½28, 29, 31 and spermathecae in 14–17................ G. singaporensis

- Wings from 17 or 18; clitellum or spermathecae different.................................................... 10

10 Intestine from 16; wings in 17, ¾18, 18–¼21, ¼22, 22; clitellum in 15–25, 26; spermathecae in 14/15–16/17... G. malayanus

- Intestine from 18; wings in 18, ½19, 19–½24, 24; clitellum in ½17, 17, 18–25, ½26, 26; spermathecae in 15–17.... G. gatesi

11 Wings in 20, 21–¼26, 26; clitellum in 17, 18–28, 29, 30; spermathecae in 13/14–16/17................ G. kotatinggi sp. n.

- Wings in 18–19; clitellum in 16, 17–23, 24; spermathecae absent................................ G. bisegmentus sp. n.