Helicopsyche pathoumthongi, Johanson & Malm, 2007

Johanson, Kjell Arne & Malm, Tobias, 2007, Three new Helicopsyche from Laos (Trichoptera: Helicopsychidae), Zootaxa 1407 (1), pp. 13-22 : 19

Helicopsyche pathoumthongi , new species

Figs. 4 , 21–30

Helicopsyche pathoumthongi , new species, has forewing R5 densely covered by blackish rod-shaped setae ( Fig. 4 ). The species is unique in the genitalia by the nearly uniformly wide, sickle-shaped primary branch of the gonocoxite, the presence of a secondary branch of the gonocoxite that is shorter than the width of the dorsal branch, and by the long basal branches that curve slightly mediad and bear 2 distal and 2 median setae. Segment X is club shaped in lateral view, with very slender lateral processes running parallel with the lateral margins. Also H. antikleia has long, slender processes on segment X, but in H. antikleia only the anterior part of the processes are attached to segment X, while in H. pathoumthongi the processes are attached to segment along their length.

Male. Cephalic setal warts small, oval; interantennal setal warts absent. Maxillary and labial palps with brushes of sensillae on dorsal face of segments; labial palp distal segment very small, about 1/4 as long as 2nd segment; maxillary palp distal segment slightly longer than basal segment, shorter than eye diameter. Antennal scape length equal to eye diameter. Tibial spurs 1, 2, 4. Forewing brownish grey with characteristic pattern of small, pale spots scattered over wing surface ( Fig. 4 ), and with short, dark setae along R5, length 4.4 mm; hind wing length 3.3 mm, with 14 hamuli; wing venation as in Fig. 23 .

Male abdomen and genitalia ( Figs.24–30 ). Sternal process VI short, oriented posteroventrally; in ventral view narrowing towards midlength, apex slightly club-shaped, densely covered by small spines. Segment IX, in lateral view, strongly expanded anteriorly, anteroventral margin nearly straight except concave near posterior corner; with sub-marginal anterior apodeme and horizontal lateral apodeme; dorsal margin nearly straight; inner margin V-shaped in dorsal view and U-shaped in ventral view; posterior margin forms 2 short, wide lobes in ventral view. Segment X straight, narrowest at mid-length; with 9 megasetae confined to clubshaped apex; 1 pair of lateral, tube-shaped processes originating from basal part of segment X, running posteriorly and attached to central segment X forming a ridge, distal-most part separated from segment X. Superior appendage long, tubular, slightly bent upwards and club-shaped at apex. Gonocoxite primary branch nearly uniformly wide in lateral view, forming a half-circle; secondary branch slightly tapering towards apex, slightly curving ventrally in lateral view and medially in ventral view, with 1 basal, 2 median, and 1 apical setae; gonocoxite basal branch short, truncated distally, with few long setae oriented posteriorly; gonocoxite basal plate long, exceeding segment IX in lateral view, anterior part nearly equally broad; in ventral view, anterior part needle-shaped. Phallus without additional sclerites, curved posteriorly at 1/3 its length and ventrally at 2/3 its length; about equally wide along its length in lateral view; in ventral view, anterior half about as wide as in lateral view; distal half about 3x broader than anterior part.

Holotype male: LAOS PDR: Luang Namtha Prov.: Nam Ha NBCA, Lakkhammai Village, Niam Leung stream, light trap, loc. 29, 21.152°N, 101.355°E [UTM 47Q 0744602, 2339873], 749 m, 30.iv.2005, Jönsson, Malm, & Viklund ( NRM, alcohol).

Paratypes: same data as for holotype — 1 male, 1 female .

Etymology. pathoumthongi , named after Mr. Bounnam Pathoumthong for his helpful assistance in the collecting of this new species.

Figures

FIGURES 2–4. Distal part of forewings of the new Helicopsyche species. 2 — H. laoensis, new species; 3 — H. vongsombathi, new species; 4 — H. pathoumthongi, new species.

FIGURES 21–22. Helicopsyche pathoumthongi, new species, holotype. 21 — head, anterolateral view; 22 — head, frontal view.

FIGURES 23–30. Helicopsyche pathoumthongi, new species, holotype. 23 — right wings; 24 — abdominal sternite VI, lateral view; 25 — process on abdominal sternite VI, ventral view; 26 — genitalia, lateral view; 27 — genitalia, dorsal view; 28 — genitalia, ventral view; 29 — phallus, lateral view; 30 — phallus, ventral view.