Isostenosmylus Krüger, 1913a

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 72-73

Isostenosmylus Krüger, 1913a

( Figs 49–50 )

Type species. Osmylus pulverulentus Gerstaecker, 1894: 166 (by monotypy).

Diagnosis. Female forecoxa with or without pedicellate setae (occasionally numerous), sometimes with preapical process; wings broadly ovoid, membrane hyaline with irregular suffusion and small dark markings along wing veins; FW costal area relatively broad basally, subcostal veinlets rarely forked; FW stem of RP short, RP originating on RA close to wing base, first branch of RP originating close to origin of RP; RP branches strongly sinuous distally in both wings; end-twigging mostly regular and very close to margin of wing; FW M fork just beyond midpoint of wing, MA and MP regularly pectinately branched; female with all FW veins regular, not incrassate; FW CuA and CuP distally strongly arched towards posterior wing margin; FW 2A very short, terminates in wing margin before origin of first branch of RP; male tergites 8 and 9 entirely fused with distinct sulcus, ectoproct apex with angular posterior processes or lobes; gonarcus with anterior apodeme present; entoprocesses curved and apically clubbed; female sternite 8 flat and plate-like, gonapophysis 9 usually with apex bilobed.

Comments. Isostenosmylus is the most species rich genus of Stenosmylinae , with 10 described species and more undescribed species in collections (Kimmins, 1 940; Ardila-Camacho & Noriega, 2014; Martins et al., 2016; Martins, unpublished data). Species of this genus are distributed in south Brazil, and along the Andes from Argentina and Bolivia to Venezuela, with many of them having an apparent predilection to montane forests. This suggest that the many species are locally endemic, with their diversification likely related to the geological uplifting of the Andes. The larvae of this genus are recorded as arboreal ( Martins et al., 2016; Winterton et al., 2017).

Included species.

I. bifurcatus Ardila-Camacho et al. in Martins et al., 2016 ( Colombia)

I. contrerasi Ardila-Camacho & Noriega, 2014 ( Colombia)

I. fasciatus Kimmins, 1940 ( Bolívia, Peru, Venezuela)

I. fusciceps Kimmins, 1940 ( Peru)

I. irroratus Ardila-Camacho et al. in Martins et al., 2016 ( Venezuela)

I. julianae Aspöck et al. in Martins et al., 2016 ( Peru)

I. morenoi Navás, 1928 ( Ecuador)

I. nigrifrons Kimmins, 1940 ( Ecuador)

I. pulverulentus ( Gerstaecker, 1894) ( Brazil, Paraguay)

I. septemtrionalandinus Ardila-Camacho & Noriega, 2014 ( Colombia)

Figures

FIGURE 49. Stenosmylinae. Isostenosmylus sp. (photo: Steve Marshall).

FIGURE 50. Stenosmylinae. Isostenosmylus fusciceps Kimmins.