Nuddsia Menon & Makarkin, 2008

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 23

Nuddsia Menon & Makarkin, 2008

( Fig. 7D )

(= Burmaleon Myskowiak et al., 2016 syn. nov. Type species. Burmaleon magnificus Myskowiak et al., 2016: 219 )

Type species. Nuddsia longiantennata Menon & Makarkin, 2008: 151 (by original designation).

Diagnosis. Wings elongate and narrow; FW RP with five branches, RP1 originating far from origin of RP, close to wing length; FW M fork at to 1/3 wing length, slightly proximal origin of RP1; MA and MP each dichotomously branched apically in both wings; FW CuA with distal fork forming distinct loop; FW CuP and HW CuA elongate pectinate branched.

Comments. Nuddsia (late Aptian) and Burmaleon syn. nov. (earliest Cenomanian) are very similar in appearance and while being located on separate continents, are relatively close in age. Indeed, the only feature to distinguish between Burmaleon and Nuddsia is the slight difference in position of the forewing medial fork. This fork is close to, but just proximal to the origin of RP 1 in Burmaleon , while in Nuddsia it is slightly more proximal. This distinction is relative and hardly significant at the generic level when compared to other genera of Gumillinae . The overall similarity of Nuddsia and Burmaleon is otherwise close enough to consider them congeneric and Burmaleon is herein synonymised with Nuddsia . Both Martins-Neto & Rodrigues (2010) and Myskowiak et al. (2016) describe the main difference between Nuddsia longiantennata and Nuddsia repatriata as being the medial fork of forewing being positioned at the base of the wing, slightly basal to the origin of RP. Yet, no other osmylid forewing has the medial fork this close to the base of the wing, although all osmylid hind wings do. The forewing and hind wing venation in Osmylidae is at times very similar, especially in Gumillinae where the costal area is not expanded in the forewing. We are sure therefore, that the forewing of N. repatriata presented in Martins-Neto & Rodrigues (2010: fig. 2) is instead a hind wing, although the placement in Nuddsia is still valid. Nuddsia is similar in wing venation to Gumilla but differs in the position of the forewing medial fork being distal to the origin of the first branch of RP.

Included species.

N. longiantennata Menon & Makarkin, 2008 ( Brazil) (Early Cretaceous)

N. repatriata Martins-Neto & Rodriguez, 2010 ( Brazil) (Early Cretaceous)

N. magnifica ( Myskowiak et al., 2016) comb. nov. ( Myanmar) (Late Cretaceous)

Figures

FIGURE 7. Gumillinae fossils. A. Allotriosmylus uniramosus Yang et al. (from Yang et al., 2010); B. Epiosmylus panfilovi Ren & Yin (from Ren & Yin, 2002); C. Enodinympha translucida Ren & Engel (forewing); D. Nuddsia longiantennata Menon & Makarkin (from Menon & Makarkin, 2008); E. Nilionympha pulchella Ren & Engel (from Ren & Engel, 2007); F. Tenuosmylus brevineurus Wang et al. (from Wang et al., 2009b) (figures not to scale).