Epiosmylus Panfilov in Dolin et al., 1980

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 20

Epiosmylus Panfilov in Dolin et al., 1980

( Fig. 7B )

Type species. Epiosmylus longicornis Panfilov in Dolin et al., 1980: 100 (by original designation).

Diagnosis. Wings narrow, subacute apically; RP with 5–6 branches; RP1 originating far from origin of RP (1/3–1/ 2 wing length); FW medial fork close to but distal to origin of RP; MA with few branches near wing margin in both wings; FW MP pectinately branched with 3–6 branches; HW MP elongate, pectinately branched; FW CuA pectinately branched (3–5 branches); FW CuP pectinately branched (3–7 branches); HW CuA elongate with widely spaced pectinately branches; HW CuP simple; FW A1 pectinate (3 widely spaced branches).

Comments. The genus is the type of the family Epiosmylidae Panfilov in Dolin et al., 1980 , which was synonymised with Gumillinae by Menon & Makarkin (2008). Epiosmylus is differentiated from most other gumilline genera by the first RP branch originating on RP at least 1/3 wing length and relatively few branches of RP (5–6). It appears closely related to Enodinympha and Tenuosmylus . The wings of these genera are relatively narrow compared to other genera (e.g., Gumilla ).

Included species.

E. longicornis Panfilov in Dolin et al., 1980 ( Kazakhstan) (Late Jurassic)

E. longus Khramov, 2017 ( Russia) (Middle Jurassic)

E. panfilovi Ren & Yin, 2002 ( China) (Middle Jurassic)

Figures

FIGURE 7. Gumillinae fossils. A. Allotriosmylus uniramosus Yang et al. (from Yang et al., 2010); B. Epiosmylus panfilovi Ren & Yin (from Ren & Yin, 2002); C. Enodinympha translucida Ren & Engel (forewing); D. Nuddsia longiantennata Menon & Makarkin (from Menon & Makarkin, 2008); E. Nilionympha pulchella Ren & Engel (from Ren & Engel, 2007); F. Tenuosmylus brevineurus Wang et al. (from Wang et al., 2009b) (figures not to scale).