Cryptocanthon rayonensis Cook, 2002

Cook, Joyce, 2002, A Revision Of The Neotropical Genus Cryptocanthon Balthasar (Coleoptera: Scarabaeidae: Scarabaeinae), The Coleopterists Bulletin (mo 1) 56, pp. 3-96 : 49-53

19. Cryptocanthon rayonensis Cook sp.nov.

( Figs. 104–109)

Material Examined. Fourteen specimens (12 males, 2 females).

Type Material. Holotype: ‘‘ MEX [ICO]: Chiapas, 8.9 km / E. Rayon, 1,500 m / 19.IX.1991, R. Anderson / cloud forest litter, 91-109’’ ( CMNC) male . Paratypes (13):

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same data as holotype ( CMNC) 5 males, 1 female ; ‘‘ Chiapas, 10 km / W. El Bosque, 1,475 m, 15.IX. / 1992, R.S. Anderson, 92-103 / pine cloud forest litter’’ ( CMNC) 5 males, 1 female, ( INBIO) 1 male.

Etymology. The epithet ‘‘rayonensis,’’ Latin adjectival form, refers to the type locality near Rayon, Chiapas.

Diagnosis. This species is distinguished by the following combination of characters: vertical surface of clypeus not foveate; pronotal hypomeron carinate; elytra strongly angulate at suture ( Fig. 107); inflexed portion of elytron with basal fovea; metathoracic wings greatly reduced; pygidium evenly, slightly convex; paramere apices roundly angulate ( Fig. 106).

Description. Holotype. Male. Total length 4.0 mm; greatest width 2.7 mm. Head. Clypeus not strongly produced anteriorly ( Fig. 105). Clypeal teeth moderately developed, widely separated, moderately reflexed. Clypeal emargination evenly rounded; vertical surface slightly, broadly V-shaped, not foveate. Apical margin of clypeus lateral to teeth nearly straight. Disc of clypeus concave, with medium-sized punctures separated by approximately one diameter. Clypeogenal suture vaguely indicated. Gena not produced, with slight incision at junction with anterior margin of clypeus. Vertex and gena with medium-sized punctures separated by one diameter or less. Pronotum. Convex medially, laterally explanate, shallow depressions laterally in posterior half; with shallow median longitudinal depression in posterior half. Anterior angles approximately right-angled. Sides anterior to lateral angles nearly straight; slightly rounded posterior to lateral angles. Posterior angles evenly rounded, not angulate, not incised. Pronotum with medium-sized, setose punctures separated by one diameter or less; setae conspicuous, arched, pale. Pronotal hypomeron convex, coarsely punctate, with posterolateral longitudinal carina curving toward the posterior angles.

Elytra. Transversely and longitudinally strongly convex; in posterior view, strongly angulate at suture ( Fig. 107); lateral carinae not strongly elevated. Each elytron dorsally with seven striae, first six indicated by two feeble wavy lines, seventh vaguely indicated along edge of dorsal carina. Elytral intervals convex; each interval with row of closely spaced very fine setose punctures on each side adjacent to striae; setae short, pale, strongly arched. Inflexed portion of elytron moderately broad, slightly sinuate apically, with scattered, fine setose punctures; basal fovea present; striae eight and nine absent, ten complete along margin of epipleuron. Epipleuron with scattered punctures in basal half.

Wings. Metathoracic wings greatly reduced. Venter. Prosternum with scattered punctures. Mesosternum with shallow, annulate punctures; punctures small medially, large laterally. Meso-metasternal suture bluntly rounded. Metasternum concave medially; disc with minute punctures in anterior third, widely spaced, small, shallow punctures posteriorly; laterally, punctures large, shallow, annulate. Legs. Profemur ventrally with medium-sized, shallow punctures. Protibia ( Fig. 104) with bilobed medial expansion in apical half; basad lobe poorly developed, subapical lobe strong, semicircular; subapical notch absent. Mesofemur and metafemur with small, elongate punctures ventrally. Metatrochanter unmodified. Metatibia slightly angulate, not crenulate; with small subapical tooth on inner margin. Abdomen. Apical sternite with irregularly spaced, medium-sized, shallow punctures. Pygidium evenly slightly convex, with medium-sized, shallow punctures separated by one diameter or more; base with median depression. Male genitalia. Parameres ( Figs. 106, 108) with apices roundly angulate, not setose.

Variation. Male: total length 3.7–4.5 mm; greatest width 2.2–2.8 mm. Female: total length 3.3–4.2 mm; greatest width 1.8–2.5 mm; clypeal teeth more strongly developed; tibiae unmodified; metasternum flat.

Bionomics. This species has been collected from cloud forest litter, at 1,475–1,500 m elevation, during the month of September.

Geographic Distribution ( Fig. 109). This species occurs in the Mexican state of

Chiapas.