Cosmodiscus Sloane, 1907

Fedorenko, D. N., 2021, Notes on the genera Aristopus, Cosmodiscus and Metabacetus (Coleoptera: Carabidae: Pterostichini: Abacetina), with description of three new species from Vietnam, Russian Entomological Journal 30 (4), pp. 413-429 : 424-425

Cosmodiscus Sloane, 1907

Sloane, 1907: 372; Andrewes, 1920: 445; 1921: 345; 1930: 131; Straneo, 1940: 211; 1982: 339; 1983: 316; Jeannel, 1948: 442; Will, 2020a: 165; Habu, 1981: 47. — Celioinkosa Straneo, 1951: 238 ; 1982: 340–341; 1995: 185 (type species: Celioschesis kivuana Burgeon, 1935 , by original designation), syn.n.

Type species: Cosmodiscus rubripictus Sloane, 1907 (by monotypy).

DIAGNOSIS. As for Aristopus except as follows: body more robust, mandibles short, laterally rounded, dorsally depressed inside a fine lateral carina, eyes hemispherical, very prominent, pronotum with entire apical bead, without basal bead (sometimes vestigial basal bead traceable outside basal sulci), elytral parascutellar seta and discal seta d2 present or not; posterior group of USS containing eight US, abdominal sternite IV without obligatory paramedian seta.

REDESCRIPTION. Body appearance and most characters as for Aristopus , except for the following few: body more robust ( Figs 21–24 , 68–70 ); head with eyes that are more prominent and thence with nearly right angle between gena and neck, mandibles peculiar in shape. Pronotum and elytra mostly subequally wide at bases, pronotum impunctate or indistinctly punctate (to densely punctate along base in at least some Afrotropical species), with lateral margin slightly reflexed and very widely explanate towards base. Elytra wide at bases, with stria 7 adjoining basal ridge at a distance from humeral angle. Legs stronger, protibia with 3–4 lateroapical spines, mesotibia with 4–5 anterolateral spiniform setae and inner setal brush consisting of 7–8 setae. Basal two protarsomeres more strongly toothed at latero-apical angles in female, inner (anterior) tooth being very large, much larger than outer one.

Median lobe of aedeagus more or less geniculate, with a distinct longitudinal median carina in the curve of ventral margin; apex small, triangular and curved ventrad. Internal sac unarmed. Left paramere widely rounded, right one less so.

Abdominal urite VIII in female as in Figs 32–33. Female genitalia and reproductive tract (examined in C. platynotus ) as for Aristopus , except only that ventral membrane of urite IX, laterotergite and gonosubcoxite are glabrous; gonocoxite is slightly different (Fig. 35): slenderer, more curved, with basolateral angle more attenuated; single ventral ensiform seta only present.

For other distinctive features see ‘Diagnosis’.

GEOGRAPHIC DISTRIBUTION. Throughout Paleotropical realm, east to Australia and southeastern Palearctics ( Japan).

HABITATS AND HABITS. Darlington [1962] mentioned that he collected two species, C. rubripictus and C. brunneus Darlington, 1962, ‘among dead leaves and debris on the ground in rain forest’ and I have taken twelve out of totally 13 specimens at light at the edges of monsoon forests.

COMMENTS. Ten species are here recognized within the genus: C. rubripictus Sloane, 1907 (Australian and Papuan), C. brunneus (Papuan) , six Oriental species, including new two (see below), C. platynotus ( Bates 1873) , C. rufolimbatus Jedlička, 1936 (= C. louwerensi Straneo, 1940 , syn.n.), C. umeralis Andrewes, 1937 , and three species from Central and West Africa (= Celioinkosa ). Darlington [1962] supposed C. rufolimbatus and C. louwerensi to be only forms of, i.e., conspecific with, C. rubripictus .

Figures

Figs 19–24. Dorsal habitus: 19 — Aristopus picturatus, ♀ from Anuradhapura env., Sri Lanka; 20 — A. humeratus sp.n., paratype ♂ from env. Pai, Thailand; 21 — Cosmodiscus platynotus, ♀ from the Bi Doup — Nui Ba Nature Reserve; 22 — C. rufoapicalis sp.n., holotype; 23 — C. rufolimbatus, holotype; 24 — C. louwerensi, paratype. Рис. 19–24. Габитус дорЗально: 19 — Aristopus picturatus, ♀ иЗ окр. Анурадхапура, Шри Ланка; 20 — A. humeratus sp.n., паратип ♂ иЗ окр. Паи, Таиланд; 21 — Cosmodiscus platynotus, ♀ иЗ Запов. Бидуп-Нуйба; 22 — C. rufoapicalis sp.n., голотип; 23 — C. rufolimbatus, голотип; 24 — C. louwerensi, паратип.

Figs 68–70. Dorsal habitus of Cosmodiscus spp.: 68 — C. sharovae sp.n., holotype; 69 — C. umeralis, holotype; 70 — C. mirei, paratype ♂. Рис. 68–70. Габитус дорЗально Cosmodiscus spp.: 68 — C. sharovae sp.n., голотип; 69 — C. umeralis, голотип; 70 — C. mirei, паратип ♂.