Prototyphis gracilis, Houart & Héros, 2008

Houart, Roland & Héros, Virginie, 2008, Muricidae (Mollusca: Gastropoda) from Fiji and Tonga, Mémoires du Muséum national d'Histoire naturelle 196, pp. 437-480 : 473-480

Prototyphis gracilis n. sp.

Figs 7B-E

TYPE MATERIAL. — Holotype (dd) MNHN 20010 .

TYPE LOCALITY. — Fiji, Somosomo Strait , 16°45’S, 179°59’E, 416 m [BORDAU 1: stn CP 1394] .

MATERIAL EXAMINED. — Only known from the holotype.

DESCRIPTION. — Shell large for genus, holotype 23.5 mm in length, triangular, smooth, lightly built. Spire high with 1¾ protoconch whorls and 6 convex, narrow teleoconch whorls. Suture impressed. Protoconch bulbous, whorls rounded, smooth, glossy; terminal lip curved, eroded.

Teleoconch axial sculpture consisting of narrow, lamellar varices. First whorl with 4 lamellae, second to last whorls with 3 narrow, wing-like varices with fairly long, open spine-like tube on shoulder. Wings narrow, fragile, with wavy growth lines on adapertural surface. Intervarical sculpture consisting of 3 narrow, rounded, low axial ridges from first to last teleoconch whorls. Other axial sculpture of growth striae. Primary and secondary spiral sculpture indistinct on early whorls; partly visible on last whorl and on abapertural side of varical wings of last whorl but still very shallow and indistinct. Other spiral sculpture consisting of numerous, narrow lirae, variable in strength.

Aperture entirely smooth within, narrow, ovate, opening into open channel in shoulder spine. Columellar lip narrow, smooth. Inner lip adherent but broken in holotype. Outer lip erect, smooth. Siphonal canal long, strongly dorsally bent at tip, open.

Shell pale fleshy-brown with faint, darker bands on abapical part of varical wings of last whorl.

REMARKS. — Prototyphis gracilis differs from the Australian species P. angasi in having a smoother and larger shell, although with one fewer teleoconch whorl. The protoconch is rounded rather than keeled or strongly keeled in P. angasi . The teleoconch whorls are less strongly shouldered, almost rounded with smoother intervarical ridges. Prototyphis gracilis has 3 intervarical axial ridges from first to last teleoconch whorls, while P. angasi has 3 ridges on the penultimate and last whorls but the early whorls have only 2 strong ridges. The spine-like tubes in P. gracilis are broader and less upwardly bent and the siphonal canal is almost twice as long as in P. angasi .

Prototyphis gracilis differs from P. eos in having rounded rather than strongly shouldered whorls, lower and smoother axial ridges, a rounded protoconch, a longer siphonal canal, a more weakly ornamented adapertural surface of the varical wings, reduced spiral sculpture, and a smooth rather than strongly denticulate outer apertural lip.

Prototyphis gracilis resembles Murex extrinodosus Sacco, 1904 [= Murex trinodosus Tate, 1888 non Bellardi, 1872]. Tate (1888: 97) compared M. trinodosus with P. angasi ; however, in the original description of the aperture, Tate described the peristome of M. trinodosus as “continuous”. Therefore we think that this Australian Pliocene species would be better placed in Ponderia .

ETYMOLOGY. — Gracilis (Latin): slender, thin.

ACKNOWLEDGEMENTS

We are most grateful to the late R. Kilias (Museum für Naturkunde, Berlin, Germany) for the loan of the holotype of Pazinotus falcatiformis (Thiele, 1925) ; to the late R. S. Houbrick (National Museum of Natural History, Smithsonian Institution, Washington D.C., USA), for the loan of the paralectotype of Murex hystricinus Dall, 1889 ; to M.G. Harasewych (National Museum of Natural History, Smithsonian Institution, Washington D.C., USA) for the photograph of the radula of Paziella pazi (Crosse, 1869) ; to A. Warén (Natural History Museum, Stockholm, Sweden) for preparation and SEM photographs of the radulae and shells; to P. Maestrati and D. Brabant (Muséum national d’Histoire naturelle, Paris) for the digital photographs and for mounting the plates, and to A. Beu (Institute of Geological & Nuclear Sciences, Lower Hutt, New Zealand) and R. H. Cowie (University of Hawaii, USA) for useful comments on the manuscript.

TABLE 2. — Muricidae from Fiji.

TABLE 3. — Muricidae from Tonga (all from the BORDAU 2 expedition) (bold: also reported from Fiji).

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