Parachorius

Tarasov, Sergei, 2017, A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data, Zootaxa 4329 (2), pp. 101-149 : 115-116

publication ID	https://doi.org/ 10.11646/zootaxa.4329.2.1
publication LSID	lsid:zoobank.org:pub:8226E27D-E3A9-481A-B51B-558643Bbb03A
DOI	https://doi.org/10.5281/zenodo.6010806
persistent identifier	https://treatment.plazi.org/id/03C187E1-AE53-B63F-DD9E-43E9FEC2039F
treatment provided by	Plazi
scientific name	Parachorius
status

Treatment

Key to females of Parachorius View in CoL

Twelve out of 19 Parachorius species cannot be reliably identified using females. Therefore, the key below contains some couplets that refer to several species at the same time. In such ambiguous cases the distribution criterion should be used to separate species. This criterion is not always reliable as many species have overlapping distributional areas.

1 Clypeus with two teeth, outer margin of each tooth not notched basally ( Fig. 4F–G View FIGURE 4 )................................2

- Clypeus with two teeth, outer margin of each tooth distinctly or slightly notched basally ( Fig. 4D–E View FIGURE 4 )...................6

2 Clypeus with two distinctly produced teeth ( Fig. 4G View FIGURE 4 )......................................................... 3

- Clypeus with two less produced teeth ( Fig. 4F View FIGURE 4 ).............................................................. 4

3 Body black; head triangularly shaped ( Fig. 21 View FIGURE 21 ); length 6.5–6.8 mm; southern China...... P. schuelkei Tarasov View in CoL new species

- Body reddish-brown, rarely dark brown; head more rounded not as distinctly triangular as in P. schuelkei View in CoL ; length 6–7 mm .............................................. P. javanus ( Boucomont, 1914) View in CoL and P. pseudojavanus Tarasov View in CoL new species

4 Elytra covered with dense contiguous and rugose punctures ( Fig. 10F View FIGURE 10 ); pronotum almost as long as elytra; wings reduced ( Fig. 10 View FIGURE 10 ); 6–7mm; northeastern India....................................................... P. globosus Arrow, 1931 View in CoL

- Elytra covered with sparse (not contiguous) punctures ( Fig. 24F View FIGURE 24 ); pronotum notably shorter than elytra; wings normally devel- oped............................................................................................... 5

5 Length 6 mm ( Fig. 12 View FIGURE 12 ); northern India.................................................. P. hookeri Arrow, 1931 View in CoL

- Length 6.5–9.0 mm. This couplet contains four species with morphologically undistinguishable females. Their identity can be, to certain extent, determined from the geographical distribution (see species descriptions for details)..................... P. newthayerae Tarasov View in CoL new species, P. maruyamai Masumoto, Ochi View in CoL , & Sakchoowong, 2012, P. thomsoni Harold, 1873 View in CoL , P. fungorum Kryzhanovsky & Medvedev, 1966 View in CoL

6 Outer margin of each clypeal tooth slightly notched basally ( Fig. 4E View FIGURE 4 ).............................................7

- Outer margin of each clypeal tooth distinctly notched basally ( Fig. 4D View FIGURE 4 ).......................................... 8

7 Clypeal teeth acute; length 5–6 mm ( Fig. 11 View FIGURE 11 ); Taiwan................................... P. gotoi ( Masumoto, 1986) View in CoL

- Clypeal teeth rounded; length 8.2–10.6 mm ( Fig. 22 View FIGURE 22 ); northern Laos.............. P. semsanganus Tarasov & Keith, 2011 View in CoL

8 Humeral area of elytron usually with a yellow spot, sometimes the yellow spot is absent and elytra uniformly colored; India.9

- Humeral area of elytron without yellow spot, elytra uniformly colored; southeastern China, Indochina, Malay Peninsula, and Sumatra ............................................................................................10

9 Species from southern and central India ( Fig. 6 View FIGURE 6 ); length 4.5–4.7 mm ................ P. asymmetricus Tarasov View in CoL new species

- Species from northeastern India (Sikkim, Assam, Meghalaya); length 3.8–5.0 mm ( Fig. 13 View FIGURE 13 )...... P. humeralis ( Arrow, 1907) View in CoL

10 Body shape rectangular and less convex ( Figs. 7D View FIGURE 7 , 8F View FIGURE 8 ); length 3.5–4.6 mm ....................................................................................... P. bolavensis Tarasov View in CoL new species and P. fukiensis ( Balthasar, 1960) View in CoL

- Body shape round and distinctly convex ( Fig. 15F View FIGURE 15 ); length 3.8–5.4 mm. This couplet comprises four species with largely overlapping distributional areas and undistinguishable females. Nevertheless, species identity can be, to certain extent, determined from the geographical distribution (see species descriptions for details)............................................ P. longipenis Tarasov View in CoL new species, P. solodovnikovi Tarasov View in CoL new species, P. nudus ( Sharp, 1875) View in CoL , and P. peninsularis ( Arrow, 1907) View in CoL