Afropyrgota mambilla V. Korneyev, 2015

Korneyev, V. A., 2015, Pyrgotid Flies Assigned To Apyrgota. Iii. Species Of Afropyrgota Gen. N. And Tylotrypes (Diptera, Pyrgotidae), Vestnik Zoologii 49 (1), pp. 25-40 : 29-30

Afropyrgota mambilla V. Korneyev , sp. n. ( fig. 3 )

M a t e r i a l. Type. Holotype ♀ Nigeria: Mambilla Plateau , 05.04.1965, montane forest, 5000 ft (H. Roberts) ( USNM).

D i a g n o s i s. This species is similar to Afropyrgota copelandi sp. n., differing by frontal vitta at least partly brown or reddish-yellow; densely and black setulose scutum; proepisternal ridge with 8–10 long setae; scutellum with three or four moderately long black setae at each side; wing 9 mm long, grey microtrichose with two darker marks; mid femur without femoral organ, densely reddish-yellow setulose instead.

D e s c r i p t i o n. Head ( fig. 3 , 2 , 3 ) yellow, frontal vitta dark brown to black, except its posterior one-fifth orange-yellow, c. 1.3 times as long as wide in posterior half; parafacial 1.8 times as wide as flagellomere 1, brown to brownish-yellow, shining, with short and fine setulae inserted into conspicuously edged alveolae; eye twice as high as wide, 1.7 times as wide as parafacial and 4.3 times as high as gena. Face shining brownish yellow, with low, but conspicuous medial carina. Occiput entirely yellow. First flagellomere entirely reddish yellow, 2.6 times as long as wide, 1.2 times as long as pedicel and 1.9 times as long as scape. Arista entirely reddish yellow, 2-segmented. Scape 1.5 times as long as wide. Palp yellow, cigar-like, 4.5 times as long as wide, and as long as first flagellomere. Labellum very large, almost as long as eye height. Lateral vertical setae (vte) indistinguishable from postoculars or absent; holotype with two pairs of medial vertical setae (vti), of them medial 1.3 times as long as lateral vti, as long as pedicel and postocellar, and ocellar setae, 1.2–1.5 times as long as long as orbital and 2–3 times as long as mesofrontal setulae (above lunule).

Thorax yellow to brownish yellow, without contrasting dark pattern. Postpronotal lobe with 12–15 setulae and one long seta; proepisternum with 18–20 setae and setulae. Scutum as long as wide, subshining yellow with faint brown areas, moderately densely covered with long black setulae. Scutellum yellow, sparsely whitish microtrichose, with three or four setae 1.2–1.4 times as long as scutellum; no setulae ( fig. 3 , 5 ). Anepisternum yellow, with brownish posterior half bearing 25–30 long setulae in its presutural part and one long seta and 10–12 irregular long setulae in its postsutural part; anepimeron with one long seta and 15–16 irregular long setulae 0.5–0.75 times as long as seta; katepisternum with one long seta and six to seven thinner and shorter setulae. Other sclerites brownish yellow, covered with short and sparse whitish microtrichia not covering underlying cuticle.

Wing ( fig. 3 , 1 ) grey microtrichose with two darker marks posterior of stigma and in apical quarter. Wing length 15 mm. Lower calypter very narrow, without cilia; upper calypter with short black fringe basally; no visible thicker cilia.

Legs ( fig. 3 , 1 ; 6 , 7 ) entirely yellow, black setulose and setose. Coxae unmodified. Fore femur ( fig. 3 , 6 ) moderately narrow, long setose on posterior surface. Mid femur ( fig. 2 , 7 ) without femoral organ, densely reddish-yellow stulose instead. Hind femur narrow, short setulose 4–5 long subapical dorsal setae and long basiventral seta.

Abdomen narrow, cylindrical, as long as thorax, with syntergite 1+2 long, 1.5 times as long as tergites 3–6 combined.

Female terminalia not dissected. Oviscape (in situ) entirely brownish yellow, shining, sparsely black setulose, uncompressed, almost as long as preabdominal tergites 1– 6 combined, with apex bearing 2 yellow hook-like, anteromesally directed lobes ( fig. 3 , 8 , 9 ). Eversible membrane not examined.

Male unknown.

Рис. 3. Afropyrgota mambilla sp. n., ♀: 1 — голотип, вид сбоку; 2–4 — голова (2 — вид сбоку, 3 — спереди, 4 — сверху); 5 — среднеспинка, вид сверху; 6 — передняя нога, вид сзади; 7 — средняя нога, спереди; 8 — основной членик яйцеклада, вид сбоку и снизу; 9 — то же, вершина, снизу.

Figures

Fig.3. Afropyrgota mambilla sp. n., ♀: 1 — holotype, lateral view; 2– 4 — head (2 — left, 3 — anterior, 4 — dorsal view); 5 — mesonotum, dorsal view; 6 — fore leg, posterior; 7 — mid leg, anterior; 8 — oviscape, lateroventral; 9 — same, apex, ventral.

Fig. 2. Afropyrgota copelandi sp. n., ♀: 1 — sternite 6 and oviscape, ventrolateral view; 2 — same, lateroventral and dorsomedial lobes, left; 3 — aculeus; 4 — vagina and ventral receptacle; 5 — spermatheca. dl — dorsal lobe; lvl — lateroventral lobe; mva — medioventral desclerotized area; st6 — sternite 6; vr — ventral receptacle. Рис. 2. Afropyrgota copelandi sp. n., ♀: 1 — 6-й стернит и основной членик яйцеклада, снизу и сбоку; 2 — то же, латеровентральная и дорсомедиальная лопасти, слева; 3 — акулеус; 4 — вагина и вентральный семяприёмник; 5 — сперматека. dl — дорсальная лопасть; lvl — латеровентральная лопасть; mva — медиовентральная десклеротизированная область; st6 — 6-й стернит; vr — вентральный семяприёмник.

Fig. 5. Afropyrgota marshalli, holotype Ơ: 1 — lateral view; 2 — labels, 3 — wing; 4 — mesonotum, dorsal view; 5 — epandrium, right view. Рис. 5. Afropyrgota marshalli, голотип Ơ: 1 — вид сбоку; 2 — этикетки, 3 — крыло; 4 — среднеспинка, вид сверху; 5 — эпандрий, вид справа.

Fig. 1. Afropyrgota copelandi sp. n., ♀: 1 — holotype, lateral view; 2, 3 — head (2 — left, 3 — anterior view); 4 — wing; 5 — mesonotum, dorsal view; 6 — fore leg, posterior; 7 — mid leg, anterior; 8 — hind leg, anterior; 9, 10 — abdomen (9 — dorsal, 10 — lateral view). fo — femoral organ. Рис. 1. Afropyrgota copelandi sp. n., ♀: 1 — голотип, вид сбоку; 2, 3 — голова (2 — слева, 3 — спереди); 4 — крыло; 5 — среднеспинка, вид сверху; 6 — передняя нога, сзади; 7 — средняя нога, спереди; 8 — задняя нога, спереди; 9, 10 — брюшко (9 — вид сверху, 10 — сбоку). fo — феморальный орган.

Fig.6. Afropyrgota ota sp. n., ♀: 1 — holotype, lateral view; 2–4 — head (2 —left, 3 — anterior, 4 — dorsal view); 5 — wing; 6 — mid leg, anterior; 7 — prosternum, anteroventral; 8 — paratype abdomen, lateral view. Рис. 6. Afropyrgota ota sp. n., ♀: 1 — голотип, вид сбоку; 2–4 — голова (2 — вид слева, 3 — спереди, 4 — сверху); 5 — крыло паратипа; 6 — средняя нога, спереди; 7 — простернум, спереди и снизу; 8 — брюшко паратипа, вид сбоку.

Fig. 7. Afropyrgota uzungwa sp. n., ♀: 1 — holotype, lateral view; 2, 3 — head (2 — left, 3 — anterior); 4 — mesonotum dorsal; 5 — wing; 6 — fore leg, posterior; 7 — mid leg, anterior; 8 — hind leg, anterior and abdomen, ventrolateral. Scale bar 5 mm. Рис. 7. Afropyrgota uzungwa sp. n., ♀: 1 — голотип, вид сбоку; 2–4 — голова (2 — вид слева, 3 — anterior); 4 — среднеспинка сверху; 5 — крыло; 6 — передняя нога, вид сзади; 7 — средняя нога, вид спереди; 8 — задняя нога, вид спереди и брюшко, вид снизу и сбоку. Масштабная линейка 5 мм.

Fig. 8. Afropyrgota uzungwa sp. n., paratype ♀: 1 — abdomen left view; 2 — oviscape (macerated) ventral; 3 — same, ventrolateral; 4 — same, apex, enlarged, ventrally; 5 — aculeus; 6 — spermatheca. Abbreviations as on fig 2. Рис. 8. Afropyrgota uzungwa sp. n., паратип ♀: 1 — брюшко, вид слева; 2 — основной членик яйцеклада (мацерированный) вид снизу; 3 — то же, вид снизу и сбоку; 4 — то же, вершина увеличено, вид снизу; 5 — акулеус; 6 — сперматека. Сокращения как на рис. 2.

Fig. 9. Afropyrgota, ♀ abdominal sternites, macerated, ventral view: 1 — A. copelandi sp. n.; 2 — A. marshalli; 3 — A. uzungwa sp. n. Рис. 9. Afropyrgota, ♀ брюшные стерниты ♀, мацерированные, вид снизу: 1 — A. copelandi sp. n.; 2 — A. marshalli; 3 — A. uzungwa sp. n.