Euconnus (Glabriconnus) andringitrae Franz, 1968

Jałoszyński, Paweł, 2018, Two species previously placed in Euconnus s. str. belong in the endemic Madagascan subgenus Glabriconnus (Coleoptera: Staphylinidae: Scydmaeninae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2), pp. 553-558 : 555-556

Euconnus (Glabriconnus) andringitrae Franz, 1968

( Figs 1, 3 , 5–8 , 13 )

Euconnus (s. str.) andringitrae Franz, 1986: 225 , Fig. 200.

Type material. HOLOTYPE: MADAGASCAR: J ( Fig. 1 ), four labels ( Fig. 3 ): “ Madagascar-Centre / Cirque Boby 2500m / Andringitra-Ambalavao / 13.I.58 R.Paulian” [creamy white, printed], “INSTITUT / SCIENTIFIQUE / MADAGASCAR” [bluish, printed], “ Euconnus / (s. str.) / andringitrae m. / det. H.Franz ” [creamy white, handwritten and printed], “Typus” [red, handwritten] ( NHMW).

Revised diagnosis. Small species, BL about 1.5 mm; frons lacking longitudinal carinae; pronotum with two deep and distinct hypomeral carinae at each side; small lateral antebasal pronotal pits connected by deep transverse groove; aedeagus with pair of suboval lateral lobes in subapical region and apical portion of median lobe not constricted.

Redescription. Male. Body ( Fig. 1 ) strongly convex, elongate with deep constrictions between head and pronotum and between pronotum and elytra, appendages long; BL 1.55 mm; cuticle glossy, not opalescent, dark reddish-brown, only tarsi and palps lighter; setae yellowish, bristles nearly as dark as cuticle.

Head in dorsal view rounded, broadest at eyes, HL 0.30 mm, HW 0.33 mm; vertex and frons confluent, lacking longitudinal carinae and weakly but distinctly convex; supraantennal tubercles weakly elevated; tempora about 2.5 times as long as eyes in dorsal view; eyes moderately large and moderately convex, coarsely faceted. Punctures on vertex and frons very fine and sparse, inconspicuous; setae on vertex and frons indiscernible; genae and postgeneae, but not tempora, with short and sparse thick bristles. Antennae moderately long, AnL 0.80 mm; antennomeres I–VII each strongly elongate, VIII–X each strongly transverse, XI slightly broader than X, slightly shorter than IX–X together, about 1.5 times as long as broad, with subconical and indistinctly pointed apex.

Pronotum in dorsal view subtrapezoidal, broadest at anterior fourth, PL 0.38 mm, PW 0.45 mm; anterior and posterior margins weakly arcuate; lateral margins strongly rounded in anterior third and nearly straight in posterior 2/3, indistinctly sinuate just in front of posterior pronotal corners; base with one pair of small and shallow lateral pits connected by deep transverse groove; sublateral carinae present but diffuse; each side of pronotum with two parallel and deep hypomeral grooves distinct on entire length. Punctures on pronotal disc not visible at magnifications up to 80×; vestiture composed of short and dense bristles distributed along anterior pronotal margin, between and below hypomeral grooves.

Elytra egg-shaped, broadest distinctly behind middle, EL 0.88 mm, EW 0.70 mm, EI 1.25; elytral base as broad as pronotal base, lacking impressions and humeral calli. Punctures on elytral disc as fine as those on pronotum; setae sparse and short, nearly recumbent, barely noticeable. Hind wings absent.

Legs moderately long and slender; protibiae rapidly bent mesad at apices and each with one small apical denticle.

Aedeagus ( Figs 5–8 ) relatively stout and large in relation to body, AeL 0.50 mm, median lobe in ventral view broadest in subapical region where it forms pair of rounded lateral lobes, apex subtriangular, not constricted and with almost evenly rounded distal margin; endophallic structures asymmetrical and complicated; parameres with broadened apices, each with two to three long apical and subapical setae and one to two short subapical setae.

Female. Unknown.

Distribution. Southern Madagascar ( Fig. 13 ).

Euconnus (Glabriconnus) berentyensis Franz, 1986 ( Figs 2, 4 , 9–12 , 13 )

Euconnus (s. str.) berentyensis Franz, 1986: 223 , fig. 199.

Type material. HOLOTYPE: MADAGASCAR: J ( Fig.2 ), four labels ( Fig. 4 ): “S-Madagaskar / Galeriewald b. Berenty / lg. H.Franz 1969” with “Mg 42” on the reverse side [white, printed; reverse handwritten], “J” [white, printed], “ Euconnus / (s. str.) / berentyensis m. / det. H.Franz ” [creamy white, handwritten and printed], “Typus” [red, handwritten] ( NHMW) . PARATYPES (2 exx.; but see remarks): 2 ♀♀, same data as for holotype except for yellow “ Paratypus ” labels ( NHMW).

Revised diagnosis. Small species, BL about 1.5 mm; frons lacking longitudinal carinae; pronotum with two deep and distinct hypomeral carinae at each side; small lateral antebasal pronotal pits connected by faint transverse groove; aedeagus lacking suboval lateral lobes in subapical region, and apical portion of median lobe deeply constricted.

Redescription. Male. Body ( Fig. 2 ) strongly convex, elongate with deep constrictions between head and pronotum and between pronotum and elytra, appendages long; BL 1.55 mm; glossy and slightly opalescent, dark brown, only tarsi and palps lighter; setae yellowish, bristles nearly as dark as cuticle.

Head in dorsal view slightly rhomboidal, broadest at eyes, HL 0.30 mm, HW 0.30 mm; vertex and frons confluent, lacking longitudinal carinae, distinctly flattened; supraantennal tubercles weakly elevated; tempora about 2.5 times as long as eyes in dorsal view; eyes moderately large and moderately convex, coarsely faceted. Punctures on vertex and frons markedly fine and sparse, inconspicuous; setae barely discernible, spare, thin and recumbent; genae and postgenae, but not tempora, with very short, dense thick bristles. Antennae relatively short, AnL 0.70 mm; antennomeres I–VII each strongly elongate, VIII–X each strongly transverse, XI slightly broader than X, distinctly shorter than IX–X together, about 1.3 times as long as broad, with subconical and indistinctly pointed apex.

Pronotum in dorsal view subtrapezoidal, broadest at anterior fourth, PL 0.38 mm, PW 0.43 mm; sides in anterior fourth slightly impressed; anterior and posterior margins weakly arcuate; lateral margins strongly rounded in anterior third and weakly rounded in posterior 2/3, not sinuate in front of posterior corners; base with one pair of small and shallow lateral pits connected by a shallow transverse groove; sublateral carinae present but diffuse; each side of pronotum with two parallel and very deep hypomeral grooves distinct on entire length, narrow area between them forming a rounded but strongly elevated carina. Punctures on pronotal disc not visible at magnifications up to 80×; vestiture composed of very sparse thin setae, which are recumbent and barely noticeable, and of short and dense bristles distributed along anterior pronotal margin, between and below hypomeral grooves.

Elytra egg-shaped, broadest distinctly behind middle, EL 0.88 mm, EW 0.70 mm, EI 1.25; elytral base as broad as pronotal base, lacking impressions and humeral calli. Punctures on elytral disc as fine as those on pronotum; setae very sparse and short, nearly recumbent, barely noticeable. Hind wings absent.

Legs moderately long and slender; protibiae rapidly bent mesad at apices and each with a small apical denticle.

Aedeagus ( Figs 9–10 ) stout and large in relation to body, AeL 0.38 mm, median lobe in ventral view broadest in subapical region where it forms pair of rounded angles, apex strongly constricted, with sinuate sides and rounded distal margin; endophallic structures asymmetrical and complicated; parameres with broadened apices, each with two to three long apical and subapical setae and two short subapical setae.

Female. Unknown (see remarks below).

Distribution. Southernmost Madagascar ( Fig. 13 ).

Remarks. The type series of this species is composed of a male holotype and two female paratypes collected in the same locality. However, the females, although undoubtedly belong to Glabriconnus , differ from the male in the shape of the pronotum, proportions of the body parts, and even in the shape of the elytra, which in the male are broadest behind middle, and in the females broadest at or even slightly in front of the middle. I regard Franz’s identification of females as doubtful and do not include measurements and characters of these two specimens in the redescription.

Figures

Figs 1–4. Dorsal habitus of male holotypes (1–2) and their original sets of labels (3–4). 1, 3 –Euconnus andringitrae Franz, 1986; 2, 4 – E. berentyensis Franz, 1986.

Figs 5–12.Aedeagus in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views.5–8 – Euconnus andringitrae Franz, 1986; 9–12 – E.berentyensis Franz, 1986.

Fig. 13. Distribution of Euconnus (Glabriconnus) species in Madagascar, with dorsal habitus and aedeagus of each species (males of E. aterrimus Franz, 1986 are unknown; E. andranomenaus Jałoszyński, 2016 and E. februarius Jałoszyński, 2016 do not differ in external appearance).