Alloxysta

Ferrer-Suay, Mar, Selfa, Jesús, Notton, David G. & Pujade-Villar, Juli, 2013, Revision of the types of species of Alloxysta described by Cameron and Fergusson (Hymenoptera: Figitidae: Charipinae) and deposited in the Natural History Museum (London), including a key to the fauna of Great Britain, European Journal of Taxonomy 53, pp. 1-27 : 21-24

A key to identify the Alloxysta species present in Great Britain is given below.

The type material of A. tscheki and A. ullrichi has not yet been studied, so they have been excluded from the key.

1 Brachypterous wings, wing not reaching further than the end of the metasoma..........................2

– Fully winged, longer than body length..........................................................................................5

2 Wings reaching the base of the metasoma; radial cell open ......... A. marshalliana ( Kieffer, 1900)

– Wings reaching the beginning of the metasoma; radial cell closed................................................3

3 Pronotal carinae present; propodeal carinae absent; male and some females brachypterous .......... ............................................................................................................. A. halterata ( Thomson, 1862)

– Pronotal carinae absent; propodeal carinae present; males and females brachypterous...............4 4 Propodeal carinae present; F1 shorter than pedicel ........................... A. brachyptera ( Hartig, 1840)

– Propodeal carinae absent; F1 longer than pedicel .............................. A. pedestris ( Curtis, 1838)

5 Radial cell closed ( Fig. 8A )............................................................................................................6

– Radial cell open ( Fig. 8B ) or partially open ( Fig. 8C )...............................................................16

6 Pronotal carinae absent ( Fig. 8D )...................................................................................................7

– Pronotal carinae present ( Fig. 8E )................................................................................................9

7 Propodeal carinae absent ( Fig. 8F ); F1 longer than pedicel, F1-F3 subequal in length .............. .................................................................................................................... A. aperta ( Hartig, 1841)

– Propodeal carinae present, defining a plate ( Fig. 8G ); F1-F3 otherwise...................................8

8 Antennae longer than body length; F1 subequal to pedicel, F1 longer than F2 and F2 subequal to F3 ................................................................................................ A. mullensis ( Cameron, 1883)

– Antennae shorter than body length; F1 shorter than pedicel, F1-F3 subequal in length ................... .................................................................................................................. A. brevis ( Thomson, 1862)

9 Propodeal carinae present, both forming a plate ( Fig. 8G ).........................................................10

– Propodeal carinae absent ( Fig. 8F )...............................................................................................12

10 F1 subequal to pedicel; flagellomeres club-shaped and with rhinaria from F4; F1 longer than F2, F2 subequal to F3; radial cell 2.0 x as long as wide ......... A. ramulifera ( Thomson, 1862)

– F1 longer than pedicel; flagellomeres club-shaped and with rhinaria from F3; proportions of flagellomeres otherwise; radial cell longer (2.4-2.7 x as long as wide)........................11

11 Female: F2 shorter than F3; male: rhinaria begin in F1, pedicel-F3 subequal and slightly curved; propodeal plate with sides slightly curved; radial cell 2.7 x as long as wide in female and 2.4 x in male ............................................................... A. pusilla ( Kieffer, 1902)

– Female: F2 subequal to F3; male: rhinaria begin in F2, pedicel-F3 not subequal, without any curved flagellomeres; propodeal plate with curved sides; radial cell 2.4 x as long as wide in both male and female ....................................................................... A. flavicornis ( Hartig, 1841)

12 Head lighter than mesosoma; F2-F4 subequal in length; radial cell 3.0 x as long as wide; propodeum without setae in the medial area where the carinae are present in other species of Alloxysta ( Fig. 8F ) .......................................................................... A. victrix ( Westwood, 1833)

– Head same color as mesosoma; F2-F4 not subequal; radial cell shorter than 3.0 x as long as wide (2.0-2.7 x as long as wide); propodeum entirely covered by setae.....................................13

13 Flagellomeres from F3 club-shaped and with rhinaria; female with F1 subequal to F2, F2 shorter or subequal to F3; male with F1-F3 not subequal. Radial cell 2.5 x as long as wide .... ........................................................................................................ A. circumscripta ( Hartig, 1841)

– Flagellomeres club-shaped and with rhinaria from another segment; different proportions between flagellomeres; radial cell longer or shorter than 2.5 x as long as wide....................................14

14 Flagellomeres from F4 club-shaped and with rhinaria; F1 longer than pedicel, F1-F3 subequal in length; radial cell 2.4 x as long as wide ......... A. halterata ( Thomson, 1862) (some ♀♀)

– Flagellomeres club-shaped and with rhinaria from another segment; F1 longer than F2; radial cell with different proportions......................................................................................................15

15 Female: flagellomeres from F3 or F4 club-shaped and with rhinaria; F2 subequal to F3. Male: F1-F3 curved; radial cell 2.7 x as long as wide ............... A. fuscicornis ( Hartig, 1841)

– Female: flagellomeres from F2 club-shaped and with rhinaria; F2 shorter than F3. Male: F1 curved; radial cell 2.0 x as long as wide ........................................ A. leunisii ( Hartig, 1841)

16 Radial cell partially open ( Fig. 8B )................................................................................................17

– Radial cell completely open ( Fig. 8C )..........................................................................................22

17 Propodeal carinae present..............................................................................................................18

– Propodeal carinae absent...............................................................................................................20

18 Propodeum with two thick, well defined carinae, reaching the base independently and with sides curved and not plate-like; Rs almost straight; F1-F3 slightly curved in male .................... ............................................................................................................. A. pleuralis ( Cameron, 1879)

– Propodeum with two thin carinae, both forming a plate; Rs curved; F1-F3 not curved in male...................................................................................................................................................19

19 Propodeal carinae forming a plate but not protruding; F1 subequal to pedicel; radial cell 2.1 x as long as wide .................................................................................... A. citripes ( Thomson, 1862)

– Propodeal carinae both forming a plate and protruding; F1 longer than pedicel; radial cell 2.3 x as long as wide ....................................................................................... A. castanea ( Hartig, 1841)

20 Female: F2-F4 not subequal in length. Male: F2 and F3 strongly curved; radial cell 3.0 x as long as wide ................................................................................. A. macrophadna ( Hartig, 1841)

– Female: F2-F4 subequal in length. Male: without any curved flagellomeres; radial cell shorter than 3.0 x as long as wide.............................................................................................................21

21 Males and females: flagellomeres from F3 club-shaped and with rhinaria, F1 longer than F2 and radial cell 2.7 x as long as wide ................................................... A. semiaperta Fergusson, 1986

– Males and females: flagellomeres from F4 club-shaped and with rhinaria, F1 subequal or slightly shorter than F2 and radial cell longer (2.9 x as long as wide) .......... A. fuscipes ( Thomson, 1862)

22 Propodeal carinae present ( Fig. 8G ).............................................................................................23

– Propodeal carinae absent ( Fig. 8F )..............................................................................................24

23 Female: F1 subequal to F2, F2 longer than F3 and F3 subequal to F4. Male: flagellomeres from F2 club-shaped and with rhinaria, F2 curved; propodeal carinae joined, forming a plate; radial cell 2.2 x as long as wide ...................................................... A. abdera Fergusson, 1986

– Female: F2-F4 subequal in length. Male: flagellomeres from F1 club-shaped and with rhinaria; propodeal carinae thin and well defined at the top and joined at the bottom; radial cell 2.6 x as long as wide ............................................................... A. pallidicornis ( Curtis, 1838)

24 F1 subequal to F2, F2 longer than F3; radial cell 3.0 x as long as wide ............................................. .......................................................................................................... A. basimacula ( Cameron, 1886)

– F1 longer than F2, F2 subequal to or shorter than F3; radial cell shorter than 3.0 x as long as wide................................................................................................................................................25

25 F2-F4 subequal in length; R1 reaching costal margin ..................... A. crassa ( Cameron, 1889)

– F2 shorter than F3, F3 subequal to F4; R1 not reaching costal margin ............................................ .................................................................................................... A. piceomaculata ( Cameron, 1883)

Figures

Fig. 8. General Charipinae features. A. Closed radial cell (Alloxysta brevis). B. Partially open radial cell (A. macrophadna). C. Open radial cell (A. medinae). D. Pronotal carinae absent (A. brevis). E. Pronotal carinae present (A. citripes). F. Propodeal carinae absent (A. victrix). G. Propodeal carinae present (A. castanea).