Alloxysta basimacula ( Cameron, 1886 ), 2013

Ferrer-Suay, Mar, Selfa, Jesús, Notton, David G. & Pujade-Villar, Juli, 2013, Revision of the types of species of Alloxysta described by Cameron and Fergusson (Hymenoptera: Figitidae: Charipinae) and deposited in the Natural History Museum (London), including a key to the fauna of Great Britain, European Journal of Taxonomy 53, pp. 1-27 : 6-8

Alloxysta basimacula ( Cameron, 1886) stat. rev.

Fig. 2

Allotria basimacula Cameron, 1886: 87 .

Allotria caledonica Cameron, 1886: 88 . New synonymy

Allotria perplexa Cameron, 1889: 58 . New synonymy

Dilyta basimacula − Kieffer 1900: 114.

Dilyta caledonica − Kieffer 1900: 114.

Dilyta perplexa − Kieffer 1900: 114.

Alloxysta (Alloxysta) basimacula − Dalla Torre & Kieffer 1902: 38.

Alloxysta (Alloxysta) caledonica − Dalla Torre & Kieffer 1902: 38.

Alloxysta (Alloxysta) perplexa − Dalla Torre & Kieffer 1902: 39.

Alloxysta perplexa − Hellén 1931: 4.

Alloxysta basimacula − Andrews 1978: 78.

Alloxysta caledonica − Andrews 1978: 78.

Type material

Lectotype of Allotria basimacula

♀, designated by Quinlan (1974: 6), with the following labels: “ Lectotype ” (round label with blue margin), “Mugdock, Glasgow, Scotland, basimacula” (handwritten), “Cameron 96 76. Mugdoch”, “ Lectotype ♀ Allotria basimacula C. det. J. Quinlan, 1973” (handwritten), “B.M. TYPE HYM. 7.120.”, “ Alloxysta basimacula ( Cameron, 1886) det. M. Ferrer-Suay 2012”.

Holotype of Allotria caledonica

♀, with the following labels: “ Holotype ” (round label with red margins), “Cameron 96 76. Mugdock, Glasgow, Scotland ”, “ caledonica Cam. ” (handwritten), “ Holotype ♀ Allotria caledonica Cameron det. J. Quinlan, 1973”, “B.M. TYPE HYM. 7.126”, “ Alloxysta basimacula ( Cameron, 1886) det. M. Ferrer- Suay 2012”.

Lectotype of Allotria perplexa

♀, designated by Quinlan (1978: 125), with the following labels: “ Lectotype ” (round label with blue margins), [illegible], “Cameron 96-76, South Bank, River Clyde, Scotland ”, “ Lectotype of Allotria perplexa Cam. ♀, det. J. Quinlan, 1977”, “ Alloxysta ♀ det. J. Quinlan”, “B.M. Type Hym. 7.170”, “ Alloxysta basimacula ( Cameron, 1886) ♀ det. M. Ferrer-Suay 2012”.

Redescription

COLORATION. Head yellowish brown, mesosoma and metasoma brown. Scape, pedicel and F1-F3 dark yellow; F4-F11 brown. Legs dark yellow and veins yellowish brown.

HEAD. Transversely ovate, smooth and shiny, wider than height in front view; with setae below, between and a few above toruli. With few scattered setae on vertex and with many setae on face. Transfacial line 1.1 x height of compound eye. Malar space 0.4 x height of compound eye.

ANTENNA. Female: 13-segmented, filiform. All antennomeres covered with sparse setae. F1-F3 thinner and smoother than remaining flagellomeres; F4-F11 club-shaped, with rhinaria. Pedicel 1.6 x as long as wide; F1 3.6 x as long as wide; F2 3.6 x as long as wide; F3 2.9 x as long as wide; F4 2.7 x as long as wide. F1 1.3 x as long as pedicel; F1 subequal to F2; F2 1.2 x as long as F3; F4 1.2 x as long as F3; F4-F11 subequal in length, width and shape ( Fig. 2D ).

MESOSOMA. Pronotum covered by abundant setae, less at central area and distolateral margins; with two long, thick, clearly visible carinae ( Fig. 2C ). Mesoscutum smooth and shiny, round in dorsal view, with scattered setae. Scutellum also smooth and shiny, with few scattered setae, abundant on apex of scutellum. Height of mesopleural triangle along anterior margin 0.7 x height of mesopleuron. Propodeum covered by abundant setae, without carinae ( Fig. 2E ).

FORE WING. Longer than body, 1.6 x as long as mesosoma and metasoma together ( Fig. 2A ). Covered with dense pubescence; marginal setae present. Completely open radial cell, 3.0 x as long as wide. R1 short and straight; Rs long and slightly curved ( Fig. 2B ).

METASOMA. Anterior part with an incomplete ring of setae, glabrous at centre, widened laterally. Metasoma smooth and shiny, T3 and T4 clearly distinguished.

Remarks

Alloxysta basimacula was synonymized with A. macrophadna ( Hartig, 1841) by Fergusson (1986). After studying the type material, it is here considered as a different and valid species. The type series of this species consists of the lectotype female and one paralectotype male. Only the lectotype has been studied here.

Alloxysta caledonica was synonymized with A. macrophadna ( Hartig, 1841) by Fergusson (1986). It is represented in BMNH only by the holotype female. After studying the holotype of A. caledonica , it was found to be the same species as A. basimacula , so these two species are synonymized here.

Alloxysta perplexa was also synonymized by Fergusson (1986) with A. macrophadna ( Hartig, 1841) . A. perplexa is represented in BMNH by the lectotype female and ten paralectotypes (8 ♂♂ and 2 ♀♀). Comments on the type series of A. perplexa were given by Quinlan (1978: 125). However, after comparing the lectotype of A. perplexa with A. macrophadna , we found that they can be differentiated by the shape of the radial cell, completely open in A. perplexa and partially open in A. macrophadna . The main features of A. perplexa are coincident with those of A. basimacula ; for this reason we establish here this new synonymy.

Figures

Fig. 2. Alloxysta basimacula (Cameron, 1886). A. Fore wing. B. Fore wing radial cell (arrow indicates that the radial cell is completely open). C. Pronotum. D.Antenna, ♀. E. Propodeum. F. Lateral habitus, ♀.