Maladera modestula ( Brenske, 1902 )

Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke & Ahrens, Dirk, 2021, A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), Zootaxa 4922 (1), pp. 1-400 : 154

Maladera modestula ( Brenske, 1902) group

Key to species of the Maladera modestula ( Brenske, 1902) group occurring in China (³³):

1 Body large (> 8.5 mm). Parameres distinctly shorter than phallobase................ Maladera rufotestacea (Moser, 1915)

1’ Body smaller (<8.0 mm). Parameres distinctly longer than phallobase........................................... 2

2 Anterior margin of labroclypeus moderately reflexed......................................................... 3

2’ Anterior margin of labroclypeus stongly reflexed............................................................ 4

3 Parameres long, distinctly longer than phallobase. Basal lobe of left paramere long, exceeding half of paramere..................................................................... Maladera jinghongensis Ahrens, Fabrizi & Liu , sp. n.

3’ Parameres distinctly shorter than phallobase. Basal lobe of left paramere very short or absent, not exceeding basal quarter of paramere.................................................................. Maladera taiwana Nomura, 1974

4 Filiform appendages, including basal lobe, longer ( Fig. 34B )...................... Maladera modestula ( Brenske, 1902)

4’ Filiform appendages, including basal lobe, shorter ( Fig. 34F )............................ Maladera vethi ( Moser, 1916)

Figures

FIGURE 34. A–D: Maladera modestula (Brenske, 1902) (Myanmar: Putao); E–H: M. vethi (Moser, 1916) (China: Guangxi: Banbaxiang); I–L: M. rufotestacea (Moser, 1915) (Vietnam: Sa Pa). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale.