Nornalupia megacephala, M, 2002

M, B., 2002, A New Genus And Species Of The Subtribe Anisodactylina From South-Western Australia (Coleoptera: Carabidae: Harpalini), Acta Zoologica Academiae Scientiarum Hungaricae 48 (3), pp. 173-179 : 177-179

Nornalupia megacephala sp. n.

( Figs 1–11 )

Type material – Holotype, male, SW Australia, W Gully Rd. , Nornalup-Walpole Nature Park,

25.I.1979, unknown collector ( ANIC). Paratypes. 10 males, 9 females, same data as holotype .

Description – Body length in male 6.2–6.6 mm, in female 6.4–7.6 mm, width 2.4–2.6 and 2.6–2.9 mm, respectively.

Body dark brown to almost black, with labrum, anterior margin of clypeus, narrow margins of pronotum and elytra usually reddish brown; palpi, antennae and legs brown, tibiae apically and femora more or less infuscated. Upperside shining, not iridescent and without metallic tinge.

Head, measured across eyes and across neck constriction, correspondingly 0.81–0.87 and 0.66–0.72 times as wide as pronotum in male, and 0.85–0.92 and 0.69–0.76 times as wide as pronotum in female. Antennae slender, with middle segments a little more than 1.5 times as long as wide, reaching only base of elytra. Dorsal punctation variable from very fine and restricted to lateral portions of head to somewhat coarse and distributed almost throughout. Dorsal microsculpture, except for very narrowarea under each eyes covered with fine isodiametric meshes, invisible even at high magnification.

Pronotum 1.40–1.54 times as wide as long, widest in anterior third just behind lateral setigerous pores, with sides rounded anteriorly and straightly or weakly sinuately covergent in posterior half ( Figs 3–4 ). Anterior and posterior margins more or less straight, latter a little narrower than former and notably narrower than elytral base between humeal angles. Apical angles small and acute, weakly protruding, very narrowly rounded at tip. Basal angles obtusangular, more or less narrowly rounded at apex. Lateral furrowes narrow anteriorly and medially, weakly widened before basal angles and usually fused there with small and shallow basal foveae forming comparatively deep united laterobasal deppressions; lateral margins at basal angles slightly reflexed. Mediobasal portion of pronoum very convex. Punctures in laterobasal depressions most coarse and often confluent. Microsculpture invisible.

Elytra widest at middle, in male 1.40–1.44 times as long as wide, 2.32–2.43 times as long and 1.17–1.27 times as wide as pronotum; in female these indices 1.46–1.54, 2.58–2.71 and 1.19–1.28, respectively. Humeri prominent, sharp at apex, without denticles. Sutural angles acutangular, narrowly rounded at apex. Basal edge rather strongly sinuate laterally and meeting lateral margin in weakly obtuse or nearly right angle. Intervals strongly narrowed before apex. Striae somewhat thin. Marginal series consisting of 6 basal and 7 apical umbilicate pores. Microsculpture visible in female only on two lateral intervals throughout and usually also in narrow basal portion of all other intervals, consisting of fine and narrowtransverse meshes, in male obliterate meshes visible sometimes only in apical portion of lateral intervals.

Wings completely reduced. Metepisterna ( Fig. 6 ) small, hardly longer than wide, strongly narrowed posteriad. Metacoxae each only with two obligatory setigerous pores, without any additional pores.

Median lobe of aedeagus arcuate, with apical portion curved dorsad (lateral aspect) and to right; terminal lamella asymmetrical, longer than wide, moderately broad, narrowly rounded at apex, with sides roundly convergent apicad (dorsal aspect). Internal sac with dorso-lateral longitudinal chain of small spines.

Etymology. The species epithet is derived from the Greek, megas (big) and cephale (head), referring to the remarkable feature of the newspecies.

Distribution – Known only from the type locality, Nornalup-Walpole Nature Park, in the south-western corner of Western Australia.

*

Acknowledgements –IcordiallythankDr. OTTÓ MERKL andDr. GYŐZŐ SZÉL (HungarianNatural History Museum, Budapest) for providing hospitality during my visit to Budapest and Dr. MARTIN BAEHR (Zoologische Staatssamlung München) for the valuable remarks concerning the manuscript. The research was supported by the grant no. 01–04–49641 from Russian Foundation for Basic Research.

Figures

Figs 1–7. Nornalupia megacephala sp. n. Head and pronotum, lateral apect (1). Apex of left mandible, frontal aspect (2). Right margin of pronotum (3–4). Habitus (5). Right metepisternum (6). La-

Figs 8–11. Nornalupia megacephala sp. n. Median lobe of aedeagus, dorsal (8) and lateral (9) aspects. Stylus, lateral aspect (10). Stylus and valvifer, ventral aspect (11). Scale: 0.5 mm