Togoperla shan Stark & Sivec, 1991

Stark, Bill P. & Sivec, Ignac, 2008, The Genus Togoperla Klapálek (Plecoptera: Perlidae), Illiesia 4 (20), pp. 208-225 : 218-220

Togoperla shan Stark & Sivec

( Figs. 44‐50 , 59‐60 )

Togoperla shan Stark & Sivec, 1991:151 . Holotype ♂ (USNM), Chiang Mai, amp. Fang , Tambon Pa Pah, Thailand

50 44

45 47 48

Material examined. Thailand: Chiang Mai, amp. Fang, tambon Papah , 18‐20 April 1975, A.E. Gordon, 14 ♂, 4 ♀ ( USNM, BPS, PMSL) .

Adult habitus. General color brown. Head with dark pigment over ocelli and forward of M‐ line. Pronotum brown with indistinct rugosities ( Fig. 44 ). Legs brown without distinct banding.

Male. Forewing length 19‐21 mm. T 5 produced into a truncate or slightly emarginate lobe; T 6‐ T 9 typical; T 5 lobe with marginal tufts of long setae ( Fig. 45 ). Dorsal aspect of hemiterga moderately long with prominent sulcus and with tips curved slightly laterad; length from basal callus to hemitergal tip about 1.6‐2.0 times as long as basal callus; lateral aspect triangular in shape but tip blunt ( Fig. 46 ); sensilla basiconica clustered on basal callus and scattered on hemitergal tip. Membranous aedeagal sac coarsely armed with a broad band of short thick spines; band ends near sac apex on dorsal surface but continues onto tube on venter ( Figs. 47‐48 ); sac apex trilobed; tube with a pair of unarmed lateral lobes.

Female. Forewing length 29‐31 mm. Subgenital plate parabolic with emarginate apex ( Fig. 49 ). Vagina constricted beyond midlength ( Fig. 50 ); accessory glands short, much less than half vaginal length.

Egg. Length 0.45‐0.47 mm, width 0.35‐0.37 mm. Outline pear shaped with wide posterior pole and narrow collar; collar width 0.12‐0.14 mm, length about 0.01‐0.02 mm. Collar distinct but sessile, rim smooth but with a row of pits circling base ( Fig. 59 ). Chorion relatively smooth but with faint follicle cell impressions ( Fig. 60 ). Micropyles subequatorial with canals slanted.

Distribution. Known only from Thailand.

Remarks. Other than T. poilanina , this is the only Togoperla species known to have a distinct egg collar but in male and female genitalic features it is typical of the genus. The eggs of T. poilanina are coarsely punctate and those of T. shan lack punctations.

Figures

Figs. 44‐50. Togoperla shan. 44. Head and pronotum, 45. Male terminalia, 46. Male hemitergum, lateral, 47. Aedeagus, lateral aspect, 48. Aedeagus, dorsal aspect, 49. Female terminalia, 50. Vagina.

Figs. 57‐62. Scanning electron micrographs of eggs. 57. T. canilimbata, 58. T. canilimbata micropyles and chorionic detail, 59. T. shan collar, 60. T. shan micropyles and chorionic detail, 61. T. tricolor, from holotype, 62. T. tricolor micropyle and chorionic detail.