Raveniola birecikensis, Zonstein & Kunt & Yağmur, 2018

Zonstein, Sergei, Kunt, Kadir B. & Yağmur, Ersen A., 2018, A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia, European Journal of Taxonomy 399, pp. 1-93 : 21-22

publication ID	https://doi.org/ 10.5852/ejt.2018.399
publication LSID	lsid:zoobank.org:pub:E836E138-D6E2-4F62-B4B3-CE2E073F2B24
DOI	https://doi.org/10.5281/zenodo.5980290
persistent identifier	https://treatment.plazi.org/id/1DB28F4C-3186-4E0E-9769-C533F62AB5D4
taxon LSID	lsid:zoobank.org:act:1DB28F4C-3186-4E0E-9769-C533F62AB5D4
treatment provided by	Plazi
scientific name	Raveniola birecikensis
status	sp. nov.

Treatment

Raveniola birecikensis View in CoL sp. nov.

urn:lsid:zoobank.org:act:1DB28F4C-3186-4E0E-9769-C533F62AB5D4

Figs 50 View Figs43–51 , 72–73 View Figs 68–73 , 106 View Figs 102–116 , 133–134 View Figs 133–136 , 153–154 View Figs 153–156 , 180–181 View Figs 180–188 , 221 View Figure221

Diagnosis

The male of this species differs from those of other members of the same group by having a welldeveloped keel in the flattened proximal part of the embolus ( Figs 180–181 View Figs 180–188 ; cf. Figs 183–188 View Figs 180–188 ). Raveniola biresecensis sp. nov. can be distinguished from R. nana sp. nov. (male of the latter species is unknown) by its noticeably larger body size, more intensely brownish coloration, and relatively smaller AMEs and PMEs ( Fig. 106 View Figs 102–116 ; cf. 122).

Etymology

The specific epithet is a toponym that refers to the locality where the paratypes were collected: Birecik District, Şanlıurfa Province of Turkey.

Material examined

Holotype TURKEY: ♂, Şanlıurfa Province, Halfeti District, Şanlıurfa, N of Halfeti , 37°16′ N, 37°51′ E, 400– 600 m, 7 Mar. 2006, A.E. Yağmur leg. ( AUZM). GoogleMaps

Paratypes TURKEY: 2 ♂♂, Şanlıurfa Province, Birecik District, Yukarı-Habip , 11 km of NNE Birecik , 37°04.2′ N, 38°00.5′ E, 550–650 m, 1 Apr. 2006, A.E. Yağmur leg. ( AUZM, TAU). GoogleMaps

Aditional material (5 possibly dwarf ♂♂)

TURKEY: 3 ♂♂, Şanlıurfa Province, Halfeti District, 37°16′ N, 37°51′ E, 400–600 m, 14 Feb. 2011, E.A. Yağmur leg. ( AUZM and TAU); GoogleMaps 2 ♂♂, surroundings of Birecik, 37°04′ N, 38°00′ E, 550–650 m, 4 Feb. 2011, E.A. Yağmur leg. ( AUZM, TAU). GoogleMaps

Description

Male (holotype)

HABITUS. See Fig. 50. View Figs43–51

MEASUREMENTS. TBL 14.77, CL 4.66, CW 4.08, LL 0.28, LW 0.66, SL 2.26, SW 2.02.

COLOUR. Carapace, palps and legs dorsally light yellowish brown, with darkened cephalic area, thoracic grooves and leg I; eye tubercle dark brown; chelicerae medium cherry-red; sternum, labium, maxillae, palps and legs ventrally light yellowish brown; abdomen dorsally light yellowish grey, with indistinct darker dorsal pattern, ventral abdominal surface and spinnerets pale yellowish grey.

PROSOMA. Carapace and chelicerae as shown in Fig. 72 View Figs 68–73 . Clypeus and eye group as in Fig. 106 View Figs 102–116 . Eye diameters and interdistances: AME 0.12(0.16), ALE 0.20, PLE 0.14, PME 0.07, AME–AME 0.19(0.16), ALE–AME 0.10(0.08), ALE–PLE 0.06, PLE–PME 0.05, PME–PME 0.44. Each cheliceral furrow with 9 promarginal teeth and 9–10 mesobasal denticles. Maxillae with 7 relatively small cuspules each.

LEGS. Tibia and metatarsus I as shown in Fig. 133 View Figs 133–136 . Scopula: distal on metatarsi I–II, entire on tarsi I–II; absent on tarsi III–IV. Trichobothria: 2 rows of 9–11 each on tibiae, 15–17 on metatarsi, 10 on tarsi I and II, 10–12 on tarsi III and IV, 7 on cymbium. Paired claws with 8–10 teeth on each margin.

LEG MEASUREMENTS.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	2.40	1.35	1.88	–	0.84	6.47
Leg I	3.98	2.29	3.04	3.07	1.63	14.01
Leg II	3.61	2.17	2.67	2.68	1.64	12.77
Leg III	2.92	1.76	1.89	2.76	1.58	10.91
Leg IV	3.66	2.16	3.27	4.30	1.74	15.13

SPINATION. Femora I–IV with one basodorsal spine and 3 distal bristles; patellae I and II aspinose. Palp: femur d1–1–1–1–1, pd1; patella p1; tibia d1–1, p1–1–1, r1, v1–1; cymbium d4(5). Leg I: femur pd0–0(1)–1; tibia rv0–1–m–m, v1–1(0)–0l; metatarsus v0–0–2. Leg II: femur pd0–0–1; tibia p0–0–1, v2–2–3; metatarsus p1, v2–2–3. Leg III: femur pd0–1–1, rd0–1–1; patella p1, r1; tibia d1–0–0, p1– 1(0)–1, r1–1–1, v2–2–3; metatarsus d0–1–0, p1–1–1, r1–1–1, v2–2–3. Leg IV: femur pd0–1–1, rd0–0– 1; patella p1, r1; tibia d1–0–0, p1–1–1, r1–1–1, v2–2–3; metatarsus d0–1–1–0, p1–1–1–1, r1–1–1–1, v2–2–2–3.

PALP. Tibia, cymbium and bulb as shown in Fig. 153 View Figs 153–156 . Wide and flattened basal part of embolus with subapical keel extending into twisted and flagelliform apical part ( Figs 180–181 View Figs 180–188 ).

SPINNERETS. PMS: length 0.49; diameter 0.13. PLS: maximal diameter 0.41; length of basal, medial and apical segments 0.67, 0.42, 0.47; total length 1.56; apical segment shortly digitiform.

Female

Unknown.

Variation

Carapace length in males of the type series varies from 3.93 to 4.66. Colouration, eye arrangement, shapes of the palpal tibia, cymbium, palpal organ, tibia and metatarsus I in the paratypes do not differ significantly from those of the holotype ( Figs 73 View Figs 68–73 , 134 View Figs 133–136 , 154 View Figs 153–156 ). The appearance and the corresponding structures of the pale-coloured and possibly dwarf conspecific males (CL 2.16–3.41), possessing larger AMEs and an invariably shorter basal portion of the embolus, are shown in Figs 47 View Figs43–51 , 74 View Figs 74–78 , 107 View Figs 102–116 , 135 View Figs 133–136 , 155 View Figs 153–156 and 182 View Figs 180–188 , respectively.

Ecology

The type series, which includes only males, was collected with pitfall traps in open steppe biotopes sparsely covered with low forest and shrubs ( Fig. 230 View Figs 230–235 ).

Distribution

The species is known from south-eastern Turkey (Şanlıurfa Province) and probably also occurs in the adjoining territory of Syria. See Fig. 221 View Figure221 .