Jenningsina aff. lethiersi Becker, 1971

Maillet, Sebastien, Milhau, Bruno, Vreulx, Michel & Posada, Luis-Carlos Sánchez De, 2016, Givetian ostracods of the Candás Formation (Asturias, North-western Spain): taxonomy, stratigraphy, palaeoecology, relationship to global events and palaeogeographical implications, Zootaxa 4068 (1), pp. 1-78 : 43

Jenningsina aff. lethiersi Becker, 1971

(Pl. III: Fig. 3 )

? 1964 Jenningsina cf. sp. F; Magne: pl. 23, figs. 144–145, pl. 24, figs. 172–174.? 1970 Jenningsina nov. sp., aff. catenulata (Van Pelt) type 1; Lethiers: 72, pl. 7, figs. 7–8.? 1970 Jenningsina nov. sp., aff. catenulata (Van Pelt) type 2; Lethiers: 72, pl. 7, figs. 9–11.? 1970 Jenningsina nov. sp., aff. catenulata (Van Pelt) type 3; Lethiers: 72, pl. 7, fig. 12. cf. 1971a Jenningsina lethiersi nov. sp.; Becker: 49–51, pl. 9, figs. 80–86.

aff. 1974 Jenningsina lethiersi Becker; Lethiers: 75, 77, figs. 1, 2.

aff. 1977 Jenningsina lethiersi Becker; Casier: 2, 4, 6, 7, 9, 12, 14, 15.

aff. 1982 Jenningsina lethiersi Becker ; Milhau: 22, pl. 5, fig. 116.

non 1983a Jenningsina lethiersi Becker ; Milhau: 351, pl. 1, fig. 2.

aff. 1985 Jenningsina lethiersi Becker ; Lethiers et al.: 76–77, pl. 13, fig. 1. aff. 2007 Jenningsina lethiersi Becker ; Casier & Préat: 296, pl. 3, figs. 7–8. aff. 2008a Jenningsina lethiersi Becker ; Casier & Olempska: 55, pl. 2, fig. 9. aff. 2008b Jenningsina lethiersi Becker ; Casier & Olempska: 646, fig. 4A.

aff. 2009 Jenningsina lethiersi Becker ; Casier in Casier & Préat: pl. 5, fig. 14. 2013 Jenningsina lethiersi Becker ; Maillet: 120, pl. XV, fig. 21.

Material. 6 specimens (4 C, 2 V).

Diagnosis. Becker (1971a, p. 50).

Description. Medium-sized elongated carapace, bean-shaped in lateral view. Overlap of the larger LV over the smaller RV pronounced postero-dorsally and very slight antero-dorsally. Hmax at 1/3 anterior; Lmax median; Wmax median. Convex dorsal margin. Almost straight ventral margin. Ventral and dorsal margins converging both anteriorly and posteriorly. Free margins of the carapace flattened. Regularly convex anterior margin in its ventral part, more arched in its dorsal part. Well rounded posterior margin, oblique anteriorly from dorsal to ventral part, more arched in its dorsal part. Posterior part of the carapace slightly higher than anterior one. Surface reticulated. Globally longitudinal reticulation, with large irregular to polygonal reticles diverging posteriorly from the median longitudinal line of the valves. Marked posterior vertical shoulder along the posterior margin on each valve.

Discussion. Morphological differences between J. paffrathensis Krömmelbein, 1954 and J. lethiersi Becker, 1971 are very tenuous. Exhibiting a large size of reticles and an outline of the carapace less oval than in J. paffrathensis Krömmelbein, 1954 , the specimens of the Candás Fm show morphology close to J. lethiersi Becker, 1971 . Nevertheless, these specimens may be synonym of Jenningsina paffrathensis Krömmelbein, 1954 (ecomorphism?).Indeed, except a difference in the size of the reticules, the general pattern of reticulation is identical in these two species. On the contrary to J. paffrathensis Krömmelbein, 1954 , J. lethiersi Becker, 1971 is not known with certainty in Upper Givetian deposits, but has been reported at the extreme base of the Frasnian both in the Ardenne (e.g. Milhau 1983a) and in the Boulonnais (e.g. Lethiers 1970, 1974).However, the two species are frequently associated within a same stratigraphical level (e.g. Maillet 2013 and Casier et al. 2013 for the Ardenne; this paper for the Asturias).

Occurrences. Upper Givetian to Lower Frasnian (Candás Fm, mbs C and D), Asturias (Peran-Perlora section).

PLATE III. Ostracods from the type-area of the carbonate Candás Formation. For samples stratigraphic position, see Figure 3. 1 . Jenningsina catenulata ( Van Pelt, 1933), II1, RV (C. O.F.L.S. 922); 2. Jenningsina paffrathensis Krömmelbein, 1954 , Q8, RV (C. O.F.L.S. 926); 3. Jenningsina aff. lethiersi Becker, 1971 , U12, RV (C. O.F.L.S. 925); 4. Quasillites (Quasillites) quasilitiformis ( Polenova, 1952) , Q8, LV (C. O.F.L.S. 938); 5–6. Quasillites (Jennillites) ambiguus Becker, 1989 , juvenile (5), A9, LV (C. O.F.L.S. 943); adult (6), I6, LV (C. O.F.L.S. 1126); 7–8. Loquitzella mesodevonica Zagora, 1968 , adult (7), A9, LV (C. O.F.L.S. 958); juvenile (8), II1, RV (C. O.F.L.S. 959); 9. Podocopina indet. P1, III6, RV? (C. O.F.L.S. 965); 10– 11. Ampuloides cf. avus Zagora, 1967 , adult (10), II1, RV (C. O.F.L.S. 977); adult (11), B10, ventral view (C. O.F.L.S. 978); 12. Microcheilinella cf. affinis Polenova, 1955 , B10, RV (C. O.F.L.S. 988); 13. Tubulibairdia cf. clava ( Kegel, 1932), A9, RV (C. O.F.L.S. 1011); 14. Tubulibairdia sp. P1, B10, RV (C. O.F.L.S. 1017); 15. Bairdiocypris symmetrica ( Kummerow, 1953) , A9, RV (C. O.F.L.S. 1022); 16. Bairdiocypris vastus Polenova, 1952 , N2, RV (C. O.F.L.S. 1026); 17. Praepilatina praepilata ( Polenova, 1960) , juvenile, A9, RV (C. O.F.L.S. 1042); 18. Saumella sp., M2, RV (C. O.F.L.S. 1044); 19. Acratia ? sp. indet., M2, RV (C. O.F.L.S. 1055).

Figures

FIGURE 3. Stratigraphical distribution of ostracod taxa through the Candás Formation in the Peran-Perlora and Carranques sections. Samples numbers are replaced on a detailed stratigraphical column of the series, on which macro- and microfaunas’ occurrences are indicated.