Tumeochrysa Needham, 1909

Wu, Jingyu & Liu, Xingyue, 2024, Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China, Zootaxa 5540 (1), pp. 1-169 : 80-81

Subgenus Tumeochrysa Needham, 1909 stat. nov.

Tumeochrysa Needham, 1909: 204 . Type species: Tumeochrysa indica Needham, 1909: 204 . Original designation and monotypy.

Chrysoplecta Navás, 1910a: 55 . Type species: Chrysoplecta immaculata Navás, 1910a: 56 . Monotypy.

Diagnosis. Body medium to large. Body length 13.0– 19.5 mm, forewing length 16.5–24.0 mm, hindwing length 14.9–22.0 mm. This subgenus is characterized by the broad head some species with reddish triangular markings on the vertex, the broad scape concaved or with tubercles, the frons, clypeus and gena not marked, the not pointed terminal palpomere of maxillary and labial palpi, the thorax with reddish stripes near middle and brown markings on lateral side, the legs with dense setae and without spots, the meso- and metatibia with a spur at tip, the brown pretarsal with basal dilation, the long forewing without shadings, the indistinct pterostigma, the triangular or quadrate cell im, the 1st rs-psm crossvein located at 2/3 or 1/2 of the cell im, the gradates in three to five divergent rows, the basal crossvein of inner gradate series not meeting Psm, the closed cell dcc, the male sternum VIII not fused with IX, the elongated male sternum IX and about twice as long as sternum VIII, the tergum IX fused with ectoproct, the median arch of gonarcus with projections and invagination, the lateral arm expanded, the broad entoprocessus posteriorly with many long gonosetae on associated membrane, the arcessus rounded at base, the apically bilobed female subgenitale, the thin spermatheca, the short vela, the absence of the ventral impression, and the long and coiled duct.

Distribution. Oriental region.

Remarks. The length and shape of antennal scape are important identification characters for species of this subgenus. The male sternum IX is obviously elongated and its length is stable, but the shape of the projections on the gonarcus median arch are more important diagnostic characters.

Key to species of Tumeochrysa in China *

1. Antennal scape with tubercles or concaved................................................................. 2 Antennal scape without tubercles or not concaved........................................................... 6

2. Antennal scape with tubercles........................................................................... 3 Antennal scape concaved on inner and outer side.................................................. T. longiscape

3. Scape with prominent tubercle only on inner side; median arch of male gonarcus with tapered projections...... T. magnifica Scape with tubercle only on outer side or both on outer and inner side; projections on median arch of male gonarcus broader 4

4. Projections on median arch of male gonarcus short and broad, invagination shallow and broad............... T. praeclara Projections on median arch of male gonarcus long and slender, invagination deep and narrow......................... 5

5. Projections on median arch of male gonarcus short and only reached at base of arcessus....................... T. sinica Projections on median arch of male gonarcus obviously long and reached at 2/3 of arcessus................... T. tibetana

6. Length of antennal scape about three times of wide; projections on median arch of male gonarcus forked at tip T.biloba sp.nov. Length of antennal scape about twice of wide; projections on median arch of male gonarcus not forked................. 7

7. Projections on median arch of male gonarcus short and broad, invagination broad.................................. 8 Projections on median arch of male gonarcus long and slender, invagination deep and narrow................... T. yunica

8. Body relatively small; cell im of forewing quadrate; invagination on median arch of male gonarcus trapezoid................................................................................................... T. minina sp. nov.

Body relatively large; cell im of forewing narrow and triangular; invagination on median arch of male gonarcus U-shaped.......................................................................................... T. yangi sp. nov.

* T. issikii and T. immaculata are not included in the key.