Scopaeus mixtus Cameron, 1941

Frisch, Johannes & Narakusumo, Raden Pramesa, 2022, Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species, Soil Organisms 95 (1), pp. 23-73 : 31-33

publication ID	https://doi.org/ 10.25674/so95iss1id311
DOI	https://doi.org/10.5281/zenodo.10724260
persistent identifier	https://treatment.plazi.org/id/03B7C77A-FFC8-FFFE-BF70-FA960FD30561
treatment provided by	Felipe
scientific name	Scopaeus mixtus Cameron, 1941
status

Treatment

Scopaeus mixtus Cameron, 1941 View in CoL

( Figs 6 View Figures 1–7 , 31 View Figures 27–34 , 62– 64 View Figures 62–67 , 123 View Figures 119–133 , 149, 151 View Figures 146–154 )

Scopaeus mixtus Cameron, 1941: 229 View in CoL , 230.

Type specimens examined: Holotype ♂, Malaysia, Sabah: Sandakan, leg. Cameron; labeled ‘Type’ (red edged, round, printed label), ‘Sandakan, / N. Borneo. / Dr. Cameron.’ (printed), ‘S. / mixtus / TYPE Cam.’ (handwritten), ‘M. Cameron / Bequest / B.M.1955- 147.’ (printed), ‘ Holotype / Scopaeus mixtus / Cameron 1930 / label by J. Frisch, 2022 ’ (red, printed) ( NHML). Paratypes: 2 ♂, 2 ♀, same labels as holotype except type label, subsequent paratype label ‘ Paratype / Scopaeus mixtus / Cameron 1930 / label by J. Frisch, 2022 ’ (red, printed) ( NHML).

The type specimen represents a holotype by original designation ( ICZN 1999 : Article 73.1.1.), because Cameron (1941: 230) stated ‘Type in my collection.’ and labeled the specimen accordingly. Four additional specimens from the Cameron collection at NHML are rated as paratypes and labeled accordingly.

New records: Malaysia, Sabah: Sandakan , S Lokan, 9.1996, 3.1997, leg. Chung ( NHML, MFNB); (04°63’50’’ - 04°77’16’’N, 117°43’83’’ - 117°70’31’’E), 300 m, 5. – 6.2012, leg. SAFE Project ( HECO, MFNB) GoogleMaps .

Redescription: Habitus and coloring as in Fig. 6 View Figures 1–7 . Head subquadrate. Antennae with penultimate segment quadrate. Mesotibia slender. Surface of head and elytra matt due to dense setose punctation; pronotum shiny with very fine, sparse punctation; forebody without microreticulation. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body reddish medium brown to dark brown except for elytra gradually blackened from about longitudinal middle towards contrasted, yellow-brown posterior fifth and yellow-brown tip of abdomen; maxillary palpi, antennae and legs yellow brown. Total body length 2.5 – 2.9 mm; forebody length 1.4 – 1.5 mm.

Male: Abdominal sternite VII with unmodified posterior margin.AbdominalsterniteVIIIwithconcaveemargination in almost posterior third ( Fig. 123 View Figures 119–133 ). Aedeagus ( Figs 31 View Figures 27–34 , 62 – 64 View Figures 62–67 ) about 0.7 mm long; apical lobes narrow, hooked ventrally with slender, emarginate apices ( Figs 31 View Figures 27–34 , 62 View Figures 62–67 ), in ventral and dorsal view ( Figs 63, 64 View Figures 62–67 ) subparallel with acute ends, together only third as wide as phallobase; dorsal lobe as long as apical lobes, strongly projecting from apical lobes ventrally, semicircularly curved dorsad, with conspicuous ventral dilatation evenly tapered towards acute end; lateral lobes subrectangular, somewhat projecting laterad, each with spacious row of few setae ( Figs 31 View Figures 27–34 , 62, 63 View Figures 62–67 ); median foramen transverse, short longitudinally, limited distally by strongly sclerotized, transverse ridge.

Female: Sperm pump with long, slender segments ( Fig. 151 View Figures 146–154 ); bursa hyaline with small, apical sclerotization ( Fig. 149 View Figures 146–154 ).

Distribution: Examined specimens of Scopaeus mixtus originate from Sabah in the Malaysian part of Borneo. I expect this species to occur in Kalimantan as well, which is why it is included in this contribution.