Campanulorchis Brieger

Vo, Doan Trung, Averyanov, Leonid, Maisak, Tatiana, Canh, Nguyen Van, Nguyen, Huu Cuong, Dang, Minh Quan, Dang, Van Son, Truong, Quang Tam & Vuong, Truong Ba, 2022, Taxonomic treatment of the genus Campanulorchis (Orchidaceae), Phytotaxa 561 (2), pp. 173-182 : 174

Campanulorchis Brieger in Brieger et al. (1996: 750).

Type: — C. globifera (Rolfe) Brieger (≡ Eria globifera Rolfe ).

4 species in Cambodia, Laos, Vietnam, and S China (Hainan) ( Fig. 1 )

Description:—Herbs epiphytic or lithophytic. Pseudobulbs 1-leaved, ovoid, globular, to broadly ovoid, pubescent when young, distant on creeping rhizome, covered by persistent bracts. Rhizome plagiotropic, stout, semiwoody. Leaves petiolate; petiole and leaf blade pubescent abaxially, almost glabrous adaxially. Peduncle erect, 1–4-flowered, arising from the apex of new pseudobulb; peduncle, rachis, pedicel and ovary densely tomentose. Flowers medium sized, widely opening or campanulate, sepals and petals free; sepals subsimilar, densely tomentose outside, glabrous inside; petals glabrous, slightly oblique. Lip 3-lobed, recurved, moveably joined to column foot apex; median lobe oblong; side lobes erect, half-ovate, slightly oblique; disc with 2–3 keels. Column simple, shortly cylindrical, wingless, at apex with broadly conoid stelidia; column foot as long as column, perpendicular to the column base, outside densely tomentose, stigma almost circular, anther cap hemi globular, finely papillose; pollinia in form of 8 half ovoid bodies connected by small amorphous viscidium. Fruit erect or ascending, narrowly obconoid, densely hairy capsule.

Key to species

1. Inflorescence normally with 2–4 flowers; peduncle 8–12 cm long; leaves below densely white tomentose.........1. C. pulverulenta

–. Inflorescence always 1-flowered; peduncle normally 1.5–3.5 cm long; leaves below glabrous .......................................................2

2. Sepals and petals uniform ochre yellow; lip not swollen on the margin; pedicel, ovary and abaxial surface of sepals brown orange tomentose..............................................................................................................................................................................2. C. thao

–. Sepals and petals white with purple nerves at base; lip swollen on the margin towards the apex; pedicel, ovary and abaxial surface of sepals silvery white hairy ...............................................................................................................................................................3

3. Leaf petiole less than 2.5 cm long................................................................................................................................ 3. C. globifera

–. Leaf petiole 6–8 cm long............................................................................................................................................... 4. C. longipes

Figures

FIGURE 1. Distribution of Campanulorchis species. A. Campanulorchis pulverulenta. B. C. thao. C. C. globifera. D. C. longipes. Map compiled by Truong Ba Vuong and L. Averyanov.