Inca clathratus clathratus ( Olivier, 1792 )

Seidel, Matthias, Arriaga-Varela, Emmanuel & Sousa, Rafael, 2018, Catalogue of the Incini with the description of the first Archedinus species from Honduras (Coleoptera: Scarabaeidae: Cetoniinae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2), pp. 389-405 : 399

Inca clathratus clathratus ( Olivier, 1792)

( Figs 5E‒F )

Cetonia clathrata Olivier, 1792: 93 [original combination]

Trichius clathratus: SCHÖNHERR (1817) : 100 [new combination]

Inca clathratus: SCHAUM (1844) : 399 [new combination]

Inca fabricii Perty, 1830 . Junior subjective synonym.

Inca fabricii Perty, 1830: 51 [original combination]. Synonymized with I. weberi by BURMEISTER & SCHAUM (1840: 379).

Ynca fabricii: LAPORTE (1840) : 158 [combination with misspelled genus name]

Inca weberi LePeletier & Serville, 1828 . Junior objective synonym.

Inca weberi LePeletier & Serville, 1828: 381 [unjustified replacement name for Cetonia ynca ]. Synonymized with I. ynca by GORY & PERCHERON (1833: 103), with I. clathratus by SCHAUM (1844: 399).

Cetonia ynca Weber, 1801 . Junior subjective synonym.

Cetonia ynca Weber, 1801: 66 [original combination].Synonymized with I. clathrathus by SCHAUM (1844: 399).

Inca ynca: GORY & PERCHERON (1833) :103 [new combination]

Types. Cetonia clathrata : Syntypes are probably in MNHN since Olivierʼs collection is deposited there. Inca fabricii: One specimen is deposited in ZSM and could be a syntype ( SCHERER 1983). Cetonia ynca and Inca weberi : Types were not found in the Zoologisches Museum in Kiel (pers. comm. by Michael Kuhlmann).

Locality records from literature. ARGENTINA: JUJUY ( DI IORIO 2013), MISIONES ( BRUCH 1911, HAYWARD 1935 –1936, DI IORIO 2013), SALTA ( DI IORIO 2013). BRAZIL: MINAS GERAIS ( PERTY 1830). COLOMBIA: VALLE DEL CAUCA (RESTREPO- GIRALDO et al. 2003). ECUADOR: BOLÍVAR ( CAMPOS 1921). FRENCH GUIANA: CAYENNE ( OLIVIER 1792). GUIANA ( BATES 1889, HOWDEN 1968). PERU ( WEBER 1801): HUANUCO ( SOUKUP 1942), SAN MARTÍN ( SOUKUP 1942).

Additional material examined. BOLIVIA: COCHABAMBA: 1 ♁ 1 ♀ ( RBINS): Bolivia, Cristal Mayu, 450 m, iv.1984, G. Lecourt leg. SANTA CRUZ: 1 ♀ ( BCRC): Bolivia, Santa Cruz, Los Volcanes Reserve, ii.2013, S. Lingafelter leg.; 1 ♁ ( CMNC): Bolivia, Santa Cruz, Andrés Ibañez, 24.v.1992, moribundo a los pies de Jasmin [= dying at the feet of Jasmin], Zeballos-Lobo leg.; 1 ♁ ( CMNC): Bolivia, Santa Cruz, Pua Ichilo, Buenavista, ii.1950, Martínez leg.; 1 ♁ ( CMNC): Bolivia, Santa Cruz, Puia. Ichilo,Buenavista,Tacú, iii.1951, Martínez leg;1♁ ( MZSP):Bolivia,Santa Cruz,Abapo, (20 km S, 19°06ʹS 63°14ʹW, 730 m), Clarke & Zamaloa leg.; 1 ♁ ( MZSP): Bolivia, tropica, Region Chapare (400 mtr.), v.1953, Dirings leg. BRAZIL: AMAPÁ: 2 ♀♀ ( MZSP):Brasil,Amapá,Macapá, (Pôrto Platon), x.1964, E. Dente leg. AMAZONAS: 1 ♀ ( BCRC): Brazil, Tefe, XI; 1 ♁ ( ZMUH):Brazil,Manaus,Rio Negro,coll.Le Moult; 1♀ ( ZMUH):Brazil, Amazone, Lago Calado, coll. 1 ♀ ( MZSP): Brasil, Amazonas, Benjamin Constant,Rio Javari, xii.1962,Dirings leg.; 1♀ ( MZSP):Amazonas, Rio Jaruá, 1901; 1♀ ( MZSP):Amazonas,Taracuá, 1937,Pe.A.Giacomo leg.; 1♀ ( MZSP):Amazonas,Teffé,A.H.Passl leg.; Gounelle;1♁ ( RBINS):Brazil, Amazonas,Tonantins, 80 m, vii.1993, M. Büche leg. BAHIA: 1♀ ( MZSP): Brasil, Baia, Ilhéus, (Fazenda São Caetano), viii.1959, Dirings leg. BRASILIA: 1 ♁ ( CEMT): Brasil, Distrito Federal, Gama, Eldorado, 07.x.2009, J.A.I. Cordeiro leg. ESPIRITO SANTO: 1 ♀ ( DZUP): Brasil, Espirito Santo, Linhares, 50 m, iii.1964, ( DZUP 273708); 1 ♁ ( DZUP): Brasil, Espirito Santo, Linhares, 50 m, ii.1964,( DZUP 273705);1♁ ( DZUP):Brasil, Espirito Santo, Linhares, 50 m, xi.1966, ( DZUP 273706); 1♁ ( DZUP): Brasil, Espirito Santo, Linhares, xi.1965,A. Maller leg., ( DZUP (273707). GOIÁS: 1 ♁ ( NMPC):Goyaz, Rio Verde; 2♁♁ ( FZBRS): Goiás, Minaçú, Serra da Mesa, 19–30.xi.1996,A. Bonaldo leg., (Col. MCH 238640, 238641); 1 ♀ ( MZSP): Goyaz, Pereira Magalhães, leg.; 1♁ ( MZSP): Brasil, Goiaz, Rio Verde; 1♀ ( MNCN):Brazil,Goyaz,Rio Verde. MARANHÃO: 1 ♁ ( CEMT): Brasil, Maranhão, Bom Jardim,REBIO-Pos.Biol.Gurupi,Arm.Luminosa Móvel, 17–27.i.2010, F. Limeira de Oliveira, M. B. Aguiar-Neto & A. A. T. Sousa legs. MATO GROSSO: 1 ♀ ( MZSP): Brasil, Mato Grosso, Barra do Tapirapé, 21–31.xii.1965, B. Malkin leg.; 1 ♀ ( DZUP): Mato Grosso, Chapada dos Guimarães, 23.xi.1983, Exc. Dep. Zool. DZUP col., (Polonoroeste), ( DZUP 273725); 1♀ ( MZSP):Mato Grosso, Posto Indigena do Xingu – Posto Diauarum, 2.xi.–11.xii.1973, G. Kloss leg. MATO GROSSO DO SUL: 1♁ 1 ♀ ( MZSP): Bata-Porã, Fazenda Primavera, 1.iii.1993, J. Leopoldo leg. MINAS GERAIS: 1 ♁ ( MZSP): Bresil (Minas), Sertão de Diamantina,Faz.Das Melancias,E.Gounelle leg., 10.ii.1902; 1♁ ( DZUP): Brasil, Minas Gerais, Rio Verde, 40 m, ii.1960, ( DZUP 273722). PARÁ: 1 ♁ ( DBPC): Brazil, Pará, Óbidos, vi.1994; 2 ♁♁ 2 ♀♀ ( MZSP): Brasil, Pará, Km 94, Rodovia Belém-Brasília, i.1963, E. Dente leg.; 1♀ ( MZSP): Miritituba,Rio Tapajós;1♁ 1♀ ( RBINS):Amazonas,Obidios, xii.1935;1 ♁ ( RBINS):Amazonas, Obidios, xi.1988. PERNAMBUCO: 1 ♁ 1♀ ( DZUP): Brasil, Pernambuco, Caruarú, 900 m, v.1971, Joaquim Lima leg., ( DZUP 273715). RONDÔNIA: 1♁ ( DZUP):Rondônia,Vilhena, 5.xii.1986, C.Elias leg., Polonoroeste, ( DZUP 273709); 1 ♀ ( DZUP): Rondônia, Vilhena, 15.xii.1986, C. Elias leg., Polonoroeste, (DZSP 273714). RORAIMA: 1 ♁ ( DZUP):Brasil,Roraima,Maracá, 20–26.vi.1988,Mirna &Mielke,( DZUP 273720); 7 ♁♁ 6 ♀♀ ( MZSP): Brasil, Roraima, Ilha de Maracá (Estação Ecológica de Maracá) 03°22.229’ N 61°26.648’W, 18–24.v. 2015, 143 m, Van Sommeren (fruta), Ponto 7, Biffi, Falaschi, Pinheiro & Riccardi les. SÃO PAULO: 2 ♁♁ ( MZSP): Brasilien, São Paulo, Presidente Epitácio, xii.1936, coll. H. Zellibor; 1 ♁ ( MZSP): Brasilien, São Paulo, Presidente Epitácio, ii.1937, B. Pohl leg. COLOMBIA: CAQUETÁ: 1 ♁ 1 ♀ ( SLCC): Caqueta, Amazonas, 12.i.2003, S. Castro leg. HUILA: 1 ♁ 1 ♀ ( MSPC): Colombia, Huila, Gigante, iv.1988, G. Schulz Vinnh./Han. VALLE DEL CAUCA: 1 ♁ 1 ♀ ( BCRC), Colombia, Valle, 45 km W of Buga, ii.1992; 1 ♀ ( DBPC):Colombia, Calima valley, 45 km W of Buga, 17.xi.1991; 2♁♁ 1 ♀ ( DBPC): Colombia, Calima valley, 45 km W of Buga, 20.ii.1992; 1♁ ( DBPC): Colombia, Buga, Calima valley, xii.1988; 1 ♀ ( DBPC): Colombia, Buga, Calima valley, vi.1989; 2 ♁♁ 1♀ ( RBINS): Colombia, Calima valley, 45 km West of Buga valley, 1200 m, 18.xi.1991, P. Bleuzen leg. ECUADOR: BOLÍVAR: 2♀♀ ( NMPC):Ecuador,Balsapamba,R.Haemsch leg.; 2 ♁♁ 2 ♀♀ ( USNM): Ecuador, Balzapamba, F. Campos leg. CAÑAR: 2 ♁♁ 2 ♀♀ ( DBPC): Ecuador, Cañar, iii.2002. LOS RÍOS: 1 ♁ ( USNM): Ecuador, Los Rios, v.1938; NAPO: 1 ♀ ( USNM): Ecuador, Napo, Tena (13 km SW), v.1977. ORELLANA: 1♁ ( DBPC):Ecuador, Coca, v.2001; 9♁♁ 9 ♀♀ ( DBPC): Ecuador, environs de Coca, v.2011. PASTAZA: 1 ♀ ( USNM): Ecuador, Pastaza, Puyo, v.1977, P. Spangler leg.; 1 ♁ ( DBPC): Ecuador, Pastaza, xi.2000. TUNGURAHUA: 1♁ 1♀ ( USNM):Ecuador,Banos, v.1931. FRENCH GUIANA: CAYENNE: 1♀ ( UNSM):French Guiana:Montagne des Chevaux, RN2, PK 22, ii.1969, H. Dalens leg.; 1 ♀ ( BCRC): French Guiana, Montagne des Chevaux, RN2, PK 22, xii.2014, S. Brule leg.; 1♁ 4 ♀♀ ( DBPC): French Guiana, Cacao, viii.2009. PERU: 2 ♀♀ ( ZMUH): East of Peru, Nypiranga. CUZCO: 1♁ 1♀ ( RBINS): Peru, Cuzco, xii.1997, Harald Schulz-Gorbsan leg. HUÁNUCO: 1 ♀ ( NMPC): Perú, Huánuco, Tingo María, ii.2001, S. & P. Pokorný leg.; 1 ♁ 1 ♀ ( MSPC): Perú, Tingo María, viii.2016, E. Picoy leg.; 1 ♁ 1 ♀ ( ARPC): Peru, Huanuco, Tingo Maria, 1993, H. Schulz leg.; 1 ♁ 1 ♀ ( DBPC): Peru, Tingo Maria, Santa Maria district, 670 m, ix.2008; 1♀ ( CMNC):Peru, Huanuco,Tingo Maria, 18.xii.1968, J. Schunke leg. JUNÍN: 1 ♁ 4 ♀♀ ( NMPC): Perú, Satipo, F. Tippmann leg.; 2♁♁ 1♀ ( RBINS):Peru,Satipo, 700m, iv.1986, G.Lecourt leg.; 1♁ 1♀ ( RBINS): Peru, Satipo, i.1992. LORETO: 1♁ ( MNCN): Peru, Montealegre, Rio Pachitea, G. Tessmann. MADRE DE DIOS: 1 ♁ ( DBPC): Peru, Madre de Dios, Povince of Manu, Manu, 545 m. SAN MARTÍN: 5♁♁ 5 ♀♀ ( DBPC):Peru, San Martin,Juanjui, xi.2009; 1♁ ( DBPC):Peru, San Martin,Alto Nieva, xi.2011;3♁♁ 1♀ ( DBPC):Peru,San Martin,Tocache, x.2010;2♁♁ ( DBPC):Peru,Moyobamba zona affluente; 1 ♁ 1♀ ( DBPC): Peru, San Martin, Naranjos, Rioja, zona affluente, xi.2010; 1 ♀ ( ZMUH): Peru, Cumbase. UCAYALI: 1 ♁ ( LGPC): Peru, Ucalayi, Atalaya, iii. 2011; 1 ♀ ( ZMUH): Peru, Ucayali, 1914.

Remarks. GORY & PERCHERON (1833) erroneously attributed Inca ynca to Fabricius, 1801. But FABRICIUS (1801) clearly attributed the authorship to WEBER (1801). BURMEISTER & SCHAUM (1840: 420) stated that they never saw Cetonia clathrata Olivier, 1792 and therefore only listed it at the end of their work. KRAJCIK (1999) listed the year of description of Inca fabricii Perty as 1834 instead of 1830, but this species was described in the first fascicle on page 51 which was published in 1830 ( BOUSQUET 2016). LEPELETIER & SERVILLE (1828) introduced the name Inca weberi for Cetonia ynca Weber, 1801 , probably to remove the ‘tautonymy’ which, for example, is not allowed in botany. SCHAUM (1844) synonymized Inca weberi LePeletier & Serville, 1828 with Inca clathratus Olivier, 1792 .

Figures

Fig 5. Habitus views of Incini species.A–C, E, G–H, J–K, M – dorsal habitus. D, F, I, L – lateral habitus. A–B – Archedinus relictus Morón & Krikken, 1990: A –♁ paratype; B –♀ paratype. C–D – Coelocratus rufipennis (Gory & Percheron, 1833), ♁ lectotype. E‒F – Inca clathratus clathratus (Olivier, 1792), ♁. G – I. clathratus sommeri Westwood, 1844, ♀. H–J – Golinca bifrons (Olivier, 1789): H–I – ♁, J –♀. K–M – Pantodinus klugi Burmeister, 1847: K–L –♁, M –♀. All scale bars = 10 mm. Images 5A–B courtesy of D. Kovac, C–D by courtesy of G. Cuccodoro, 5E‒M: by courtesy of R. Limoges.