Tonsilla jiugongensis, Zhang & Irfan & Wang & Zhang, 2022

Zhang, Meng, Irfan, Muhammad, Wang, Lu-Yu & Zhang, Zhi-Sheng, 2022, Six new species of Tonsilla Wang & Yin, 1992 (Araneae: Agelenidae) from southern China, with the first description of the male of T. yanlingensis (Zhang, Yin & Kim, 2000), Zootaxa 5091 (2), pp. 357-372 : 360-362

Tonsilla jiugongensis sp. nov.

Figures 3A–E , 9C–D , 10

Type material. Holotype ♂: CHINA, Hubei Province, Jiugong Mountain, Yunzhong Lake , 29.40827913°N, 114.66505218°E, 1230 m, 30. VI.2020, matured on 17.IX.2020, Y. Zhong, Q.L. Lu, H. Dong & J.W. Zheng leg. (SWUC-T-AG-50-01) ; Paratypes: 1 ♂ and 3 ♀, same data as holotype (SWUC-T-AG-50-02~05) .

Etymology. The specific epithet refers to the type locality; adjective.

Diagnosis. The male of T. jiugongensis sp. nov. can be distinguished from all other congeners by the following characters: apex of conductor with two arms; the upper arm longer than wide and slightly grooved in ventral view; lower arm arc-shaped in retrolateral view ( Fig. 3B, C ). Patellar apophysis almost C-shaped and extending 1/3 of above the tibia in retrolateral view ( Fig. 3C ). The female resembles T. tautispinus ( Wang et al. 1990) in having similar atrial margins and the outline of epigyne ( Fig. 3D, E ; Zhu et al. 2017, fig. 360A), but can be distinguished by the following characters: the distance between epigynal teeth seems C-shaped in T. jiugongensis sp. nov. ( Fig. 3D ), vs. U-shaped in T. tautispinus ( Zhu et al. 2017, fig. 360A). Spermathecae pear-shaped in T. jiugongensis sp. nov. ( Fig. 3E ), vs. globular in T. tautispinus ( Zhu et al. 2017, fig. 360B).

Description. Male (holotype, Fig. 9C ). Total length 12.07. Carapace 6.38 long, 4.15 wide; opisthosoma 5.73 long, 3.45 wide. Eye sizes and interdistances: AME 0.21, ALE 0.25, PME 0.24, PLE 0.27; AME–AME 0.16, AME– ALE 0.12, PME–PME 0.13, PME–PLE 0.29, ALE–PLE 0.09. MOA 0.69 long, front width 0.52, back width 0.62. Clypeus height 0.27. Leg measurements: I 18.15 (4.75, 5.86, 4.64, 2.90); II 16.02 (4.27, 5.07, 4.10, 2.58); III 13.98 (3.80, 4.34, 3.72, 2.12); IV 18.09 (4.81, 5.69, 5.04, 2.55). Spination of legs: femur I 210, II 410, III 500, IV 320; patella I-IV 010; tibia I 016, II 026, III 016, IV 046; metatarsus I 024, II 044, III 026, IV 166.

Palp ( Fig. 3A–C ). Patella short; patellar apophysis long, almost C-shaped, extending above the 1/3 of the tibia in retrolateral view. Retrolateral and lateral tibial apophysis distinct with blunt end. Cymbium approximately 3 times longer than wide; cymbial furrow extensively sclerotized, about half of the cymbium length. Median apophysis spoon-like. Apex of conductor with two arms; the upper arm longer than wide and slightly grooved in ventral view; lower arm arc-shaped in retrolateral view. Conductor lamella indistinct. Conductor’s dorsal apophysis long, rodshaped with pointed end. Embolus filiform, originating at the 7 o’clock position.

Female (one of the paratypes, Fig. 9D ). Total length 12.37. Carapace 6.78 long, 4.28 wide; opisthosoma 6.02 long, 3.88 wide. Eye sizes and interdistances: AME 0.21, ALE 0.31, PME 0.25, PLE 0.27; AME–AME 0.21, AME– ALE 0.18, PME–PME 0.19, PME–PLE 0.38, ALE–PLE 0.11. MOA 0.69 long, front width 0.62, back width 0.74. Clypeus height 0.23. Leg measurements: I 15.31 (4.45, 5.13, 3.52, 2.21); II 12.82 (3.82, 4.21, 2.96, 1.83); III 11.54 (3.29, 3.51, 3.08, 1.66); IV 16.27 (4.71, 5.41, 4.25, 1.90). Spination: legs: femur I 220, II 310, III 400, IV 210; patella I-IV 000; tibia I-II 026, III-IV 046; metatarsus I 045, II 046, III 066, IV 086.

Epigyne ( Fig. 3D, E ). Epigynal plate longer than wide, teeth short and located anteriorly far from atrium, separated with narrow distance from each other. Atrial margin distinct, semicircular, posterior margin round. Epigynal hoods shallow, distinct, located ventro-mesally. Copulatory opening present dorso-laterally within atrium. Copulatory ducts short, tubular. Spermathecae pear-shaped, separated by a distance almost equal to their length.

Distribution. China (Hubei) ( Fig. 10 ).

Figures

FIGURE 3. Tonsilla jiugongensis sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: At, atrium; CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae.

FIGURE 9. Habitus of Tonsilla spp. A–B, male holotype and female paratype of T. jinyunensis sp. nov.; C–D, male holotype and female paratype of T. jiugongensis sp. nov.; E–F, male holotype and female paratype of T. rutunda sp. nov.; G–H, male holotype and female paratype of T. subrostrum sp. nov.; I–J, male holotype and female paratype of T. subtruculenta sp. nov.; K–L, male and female of T. yanlingensis (Zhang, Yin & Kim, 2000); M, male holotype of T. yueliangensis sp. nov.

FIGURE 10. Distribution of Tonsilla species treated here.