Microgramma squamulosa (Kaulf.) de la Sota, Opera Lilloana

Smith, Alan R., Kessler, Michael, León, Blanca, Almeida, Thaís Elias, Jiménez-Pérez, Iván & Lehnert, Marcus, 2018, Prodromus of a fern flora for Bolivia. XL. Polypodiaceae, Phytotaxa 354 (1), pp. 448-450 : 448-450

Microgramma squamulosa (Kaulf.) de la Sota, Opera Lilloana 5: 59. 1961.

= Polypodium squamulosum Kaulf., Enum. Fil. 89. 1824.

= Microgramma mortoniana de la Sota, Amer. Fern J. 63(3): 64. 1973.

Range: —Andes from southern Peru to Bolivia (BE, CH, CO, LP, SC, TA); southeastern Brazil, eastern Paraguay, northeastern Argentina, and Uruguay.

Ecology: —Very common; epiphytic and saxicolous in semihumid forests; (250) 1000–2900 m.

Notes: —This species can be recognized by its venation patterns, the long-ciliate rhizome scales, and lanceolate scales present at least in the proximal third of midvein (may be absent in old fronds). Microgramma mortoniana , recognized as a distinct taxon based mainly on the morphology of the fronds and the rhizome scales, is considered here a synonym of M. squamulosa . Through the range of this species, there is variation in the rhizome scale margins, from sparsely to densely ciliate, as well as in the shape of fertile and sterile fronds; these variations, however are not correlated.