Ceradenia L.E.Bishop, Amer. Fern J.

Smith, Alan R., Kessler, Michael, León, Blanca, Almeida, Thaís Elias, Jiménez-Pérez, Iván & Lehnert, Marcus, 2018, Prodromus of a fern flora for Bolivia. XL. Polypodiaceae, Phytotaxa 354 (1), pp. 448-450 : 448-450

Ceradenia L.E.Bishop, Amer. Fern J. 78(1): 2(-4), f. 1. 1988.

Ceradenia is characterized by lobed to pinnate (or rarely pinnate-pinnatifid) blades, lacking or with obscure hydathodes, and with conspicuous, whitish, spherical, waxlike paraphyses in the sori that are most easily seen on young leaves ( Bishop 1988). Enterosora and Zygophlebia differ by lacking paraphyses or by having brownish paraphyses that are not waxlike. Two subgenera are distinguished in Ceradenia . Subgenus Ceradenia has radially symmetric rhizomes, whitish, waxlike, glandular hairs (resembling the paraphyses) on the blade surfaces, and the petioles are short or lacking (less than half the blade length). Subgenus Filicipecten L.E.Bishop has dorsiventral rhizomes, lacks waxlike, glandular hairs on the blades, and the petioles are 1/4–3 times as long as the blades. Ceradenia is a largely neotropical genus of about 70 species, many of which are poorly known; 24 species are known from Bolivia, and 10 are endemic. Eight species are known from Africa and offshore islands ( Parris 2002). Bishop (1989a) described additional species in subg. Ceradenia , several of which occur in Peru and may eventually be found in Bolivia, such as C. dendrodoxa L.E.Bishop , C. praeclara L.E.Bishop , and C. terrestris L.E.Bishop. León-Parra (2000 , 2012) treated the genus for Colombia, recognizing 24 species, nine of which also occur in Bolivia; Labiak & Prado (2003) treated the Brazilian species of Ceradenia , three of which are present in the country.

A recent phylogenetic analysis that included both African/Madagascan and neotropical species of Ceradenia shows that the genus is monophyletic, and sister to a clade comprising Zygophlebia + Enterosora , in part ( Bauret et al. 2017).