Calapnita bario, Bernhard A. Huber, 2017

Bernhard A. Huber, 2017, Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae), Zootaxa 4219 (1), pp. 1-63 : 32-35

Calapnita bario sp. nov. Figs 120–121 , 134–141

Diagnosis. Distinguished from other species of vermiformis group by very long procursus ( Figs 134 , 288 ); from most species (except C. bariengi ) also by distal cheliceral apophyses not bipartite ( Fig. 136 ); from most (except C. dayak ) also by long and slender palpal tibia (tibia length/width>2.5; Fig. 286 ); from several species also by narrow and distinct epigynal ‘knob’ ( Fig. 141 ), and by long internal female genitalia with oval pore plates converging anteriorly and close together ( Fig. 140 ).

Etymology. The species name is derived from the type locality; noun in apposition.

Material examined. Holotype.

MALAYSIA-BORNEO: ♂, ZFMK ( Ar 16021), Sarawak, Bario , forest along river N of town (3.765– 3.771°N, 115.444– 115.448°E), 1170–1250 m a.s.l., 29.vii.2014 (B.A. Huber, S.B. Huber).

Other material. MALAYSIA-BORNEO: 1♂ 3♀ 4 juvs, ZFMK (Ar 16022), same data as holotype ; 1♀ 2 juvs in absolute ethanol, ZFMK (Bor 234), same data .

Description. Male (holotype)

MEASUREMENTS. Total body length 5.2, carapace width 0.75. Leg 1: 31.8 (7.8 + 0.4 + 7.2 + 13.6 + 2.8), tibia 2: 5.8, tibia 3: 3.6, tibia 4: 6.2; tibia 1 L/d: 103. Distance PME-PME 200 µm, diameter PME 110 µm, distance PME- ALE ~30 µm; no trace of AME.

COLOR. Entire animal pale gray to whitish, legs slightly yellowish with brown patellae and tibia-metatarsus joints.

BODY. Habitus as in Fig. 120 ; ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum slightly wider than long (0.52/0.46), unmodified.

CHELICERAE. As in Figs 136–137 , apophyses near lamellae not bipartite, with pair of lateral processes; without modified hairs; without stridulatory ridges.

PALPS. In general similar to C. vermiformis (cf. figs 139 and 140 in Huber 2011); trochanter apophysis as in Fig. 138 ; femur as in Fig. 139 , ventral processes very widely spaced (distal process at 53% of femur length; cf. Fig. 285 ); tibia length/diameter 0.92/0.33; procursus as in Figs 134–135 , very long, with simple ventro-distal flat sclerite; bulb length 0.49; embolus length 1.16; embolus tip with transparent fringes.

LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 1.5%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments barely visible in dissecting microscope.

Male (variation). Tibia 1 in other male: 7.4.

Female. In general similar to male; eye triads slightly closer together (distance PME-PME 180 µm). Tibia 1 in 3 females: 6.1, 6.1, 6.2. Epigynum as in Fig. 141 , weakly sclerotized triangular plate with small and slender anterior ‘knob’; internal genitalia as in Fig. 140 , very long, with pore plates close together and far from posterior margin.

Distribution. Known from type locality in eastern Sarawak only ( Fig. 283 ).

Figures

FIGURE 283. Known distribution of the Calapnita vermiformis group, part 1 (cf. Fig. 284). The Java record of C. lawangan is based on specimens assigned tentatively.

FIGURES 120 – 127. Live specimens, Calapnita vermiformis group. 120 – 121. C. bario sp. nov., ♂ and ♀ with egg-sac from Bario. 122 – 123. C. magaseng sp. nov., ♂ and ♀ with egg-sac from Mount Penrissen. 124 – 125. C. saluang Huber, ♂ from Berastagi and ♀ with egg-sac from Phanom Bencha. 126 – 127. C. vermiformis Simon, ♂♂ from Mount Isarog and Mount Banahaw.

FIGURES 134 – 141. Calapnita bario sp. nov. (ZFMK, Ar 16022). 134 – 135. Left procursus, prolateral and retrolateral views (arrow points at ventro-distal sclerite). 136 – 137. Right male chelicera, frontal and lateral views. 138. Left palpal trochanter, retrolateral view. 139. Left palpal femur, prolateral view. 140. Cleared female genitalia, dorsal view. 141. Epigynum, ventral view (arrow points at ‘ knob’). Scale lines: 0.5 mm (134 – 135), 0.3 mm (136 – 141).

FIGURES 285 – 288. Interspecific variation in proportions (285 – 286) and absolute sizes (287 – 288) of selected characters in the Calapnita vermiformis group. These graphs are based on measurements of holotypes and ignore intraspecific variation. The graphs show the conspicuous but largely continuous interspecific variation in some male genital characters.