Paleosisyra, Nel & Menier & Waller & Hodebert & Ploëg, 2003

Nel, André, Menier, Jean-Jacques, Waller, Alain, Hodebert, Gilbert & Ploëg, Gaël De, 2003, New fossil spongilla-flies from the lowermost Eocene amber of France (Insecta, Neuroptera, Sisyridae), Geodiversitas 25 (1), pp. 109-117 : 111

Genus Paleosisyra n. gen.

TYPE SPECIES. — Paleosisyra eocenica n. gen., n. sp.

ETYMOLOGY. — After Paleo and the genus Sisyra .

DIAGNOSIS. — Paleosisyra n. gen. has three characters absent in all extant Sisyridae , i.e. very short fusion of MA with RP in hindwing; prothoracic epimeron and episternum well defined and broad, with a furrow between them; two strong spurs on prothoracic legs (unlike the unique spur of the legs of the extant Sisyridae , after Parfin & Gurney 1956). It has also a unique combination of characters: last segment of labial and maxillary palpi triangular; mesonotum with scutellum having a broad apex; Sc and R clearly coalescent at apex of wing below the pterostigma in fore- and hindwing; costal cross-veins all simple, numerous in fore- and hindwing; series of outer and inner gradate crossveins in fore- and hindwing; anal veins A3 distally fused with A2, not running to anal margin; in male genitalia, a pair of long movable heavily sclerotized claspers, furnished with setae and denticles in their inner side; male parameres very small compared to claspers; female genital gl long and narrow, with the apex strongly curved.