Bellardiella (Szekeresia) saparuana, Varga & Páll-Gergely, 2017

Varga, András & Páll-Gergely, Barna, 2017, A review of Bellardiella Tapparone-Canefri, 1883, with descriptions of a new subgenus and two new species (Gastropoda: Cyclophoroidea: Pupinidae), Raffles Bulletin of Zoology 65, pp. 386-394 : 392-393

Bellardiella (Szekeresia) saparuana new species

( Fig. 3 L–P)

Type material. Holotype ( NHMUK 1910.12 .30.70, H = 10.6 mm, D = 5.1 mm), Saparua Ids ; Paratype ( NHMUK 1910.12.30.71), Saparua Ids .; Paratype ( NHMUK 20170015 ), Saparua Ids, coll. E.R. Sykes 1954, Acc. 1825 .

Diagnosis. A small (H: 9.1–10.6 mm, D: 4.8–5.2 mm) Bellardiella (Szekeresia) species with an elongate ovoid, very finely striated shell, open, but relatively narrow umbilicus, rounded aperture, which is slightly oblique to the shell axis in lateral view, a strong parietal callus, a strongly thickened peristome, an umbilical tube, which is attached to the peristome, and a short angular tube. Outer side of operculum multispiral without elevated lamina ( Fig. 3L ).

Description. Shell reddish brown, moderately glossy, elongate ovoid with pointed apex; the increase of the whorls is rather regular, slow at the beginning, and increasing in the last approx. 2 whorls; there are 5.5–6 whorls out of which the protoconch consists of approx. 1.5–1.75 whorls; first whorl of protoconch smooth, later with some fine radial lines; teleoconch also with fine radial riblets; sculpture of comparable strength on entire teleoconch; upper whorls moderately bulging, last two whorls less bulging; last approx. quarter whorl has a slight subsutural furrow ( Fig. 3O ); aperture rounded, sinulus absent; plane of the aperture oblique to shell axis in lateral view, with the base of the aperture protruding ( Fig. 3N ); peristome continuous with strong parietal callus; peristome strongly thickened and reflected; inner peristome is reflected; boundary between inner and outer peristomes visible in lateral and dorsal views; umbilical tube short, does not extend beyond peristome, its route visible as a keel around umbilicus, and its inner end visible by oblique view through the aperture; angular tube short, hardly projecting beyond peristome, its inner end visible by oblique view through the aperture; umbilicus open, rather narrow, deep, shows the last whorl only.

Operculum. Outer side of operculum could be observed, it is horny, glossy, multispiral, without elevated lamina.

Measurements. H = 9.1–10.6 mm, D = 4.8–5.2 mm, AH = 3.5–3.8 mm, AW = 3.9–4.3 mm.

Differential diagnosis. Bellardiella (Szekeresia) saparuana new species is most similar in shape to B. (S.) ceramica , which is more slender than the new species, has a thinner peristome, a longer angular tube, the plane of the aperture parallel to the shell axis, and elevated lamina on the outer surface of the operculum.

Type locality. Saparua Ids.

Etymology. This new species is named after the island (Saparua) where it occurs.

Figures

Fig. 3. Shells of Bellardiella (Szekeresia) species. A–E, Bellardiella (Szekeresia) lissochila Benthem-Jutting, 1963 (holotype in the NHMB); F–K, Bellardiella (Szekeresia) ceramica (von Martens, 1864) (NHMUK 20170142); L–P, Bellardiella (Szekeresia) saparuana new species (holotype, NHMUK 1910.12.30.70). Photos: A. Hänggi (A–E) and B. Páll-Gergely (F–P). Scale represents 5 mm and refers to all figures except for J, K and L.