Hypselosyrphus ulopodus (Hull, 1944)

Reemer, Menno, 2013, Taxonomic exploration of Neotropical Microdontinae (Diptera: Syrphidae) mimicking stingless bees, Zootaxa 3697 (1), pp. 1-88 : 37-38

Hypselosyrphus ulopodus (Hull, 1944)

Figs 100–104, 112.

Ubristes ulopodus Hull, 1944: 34 .

Studied type specimens. HOLOTYPE. PERU. Female. Label 1: "Lachoruria, Putmayo Distr. PERU. 17-20 Aug.

1920"; label 2: "Cornell Univ. Expedition. Lot 569"; label 3 (red): " Holotype ulopodus Hull "; label 4: " Holotype Ubristes ulopodus Hull "; label 5 (red): " Holotype Cornell U. No. 2196". Coll. CU.

Additionally studied specimens. PARAGUAY. 1 male. Label 1: " Paraguay, Vezenyi"; label 2: "Asuncion, 1904. x.5 "; label 3: "Coll. Mus. Nat. Hung."; label 4: " Ceratophya analis Curr. male, det. v. Doesburg". Coll. RMNH.

Redescription (based on holotype)

Adult female Body size: 8.5 mm.

Head. Face occupying 1/3 of head width in frontal view; yellow, except narrowly black laterally on dorsal half and with vague brown median vitta on ventral half; yellow pilose laterally, black pilose medially, with pile longer and denser around oral margin; with narrow lateral strips of white pollinosity along eye margins. Gena brownish. Oral cavity with lateral margins a slightly produced and anterior margin slightly notched. Frons black, except yellow posterior to lunula; black pilose. Vertex black; black pilose. Occiput black; black pilose dorsally, white pilose ventrally. Eye entirely with dense, dark pile, a little longer than ommati diameter. Antennal fossa about as wide as high. Antenna dark brown; antennal ratio approximately as 4:1:3.5; basoflagellomere parallel-sided with narrowly rounded apex, with small sensory pit located at 2/5 from base. Arista slender, slightly shorter than length of basoflagellomere.

Thorax. Blackish brown. Scutum entirely with long, erect black pile; with two submedian and two lateral vittae of greyish pollinosity. Postpronotum and postalar callus black pilose. Scutellum semicircular, without calcars; long black pilose anteriorly, long yellow pilose posteriorly. Anepisternum a little convex, with very slight sulcus between anterior and posterior part; black pilose, with large bare medioventral area. Anepimeron entirely black pilose. Katepimeron pilose. Katepisternum black pilose dorsally, bare ventrally. Katatergum and anatergum long and short microtrichose, respectively. Other pleurae bare. Calypter and halter dark greyish brown.

Wing: hyaline with yellow veins; tinged with brown in costal cells. Microtrichose, except for basal 1/4 of cell br and basal 1/3 of cell bm.

Legs, including coxae and trochanters, blackish brown, except apical four tarsomeres brownish yellow; entirely with long, black pile.

Abdomen. Brown. Tergites 3-5 fused, sutures not visible. Tergites black pilose laterally, pale yellow pilose medially, except tergite 4 entirely black pilose. All sternites with long black pile.

Male. (based on 1 specimen in coll. RMNH) Differs from female in the following: face black; antennal ratio approximately 4:1:2.5; tarsi entirely brownish yellow; abdomen blackish brown with posterior margin of tergite 4 yellow. Genitalia as in fig. 112.

Diagnosis. This is the only known species of Hypselosyrphus with a pilose katepimeron and a ventrally widened occiput.

Notes. This species is aberrant from its congenerics because of the pilose katepimeron, the ventrally widened occiput and the unproduced vertex (the latter character is only shared with H. pseudorhoga spec. nov.). A phylogenetic analysis based on morphological characters placed the species as a sister of the clade containing Rhoga and Hypselosyrphus (Reemer and Ståhls 2013b). The relationships within this clade are considered not well enough established to introduce another generic name, hence the current assignment of this species.

Distribution. Known from Peru and Paraguay.