Galathea squamea Baba, 1979

Macpherson, Enrique & Robainas-Barcia, Aymee, 2015, Species of the genus Galathea Fabricius, 1793 (Crustacea, Decapoda, Galatheidae) from the Indian and Pacific Oceans, with descriptions of 92 new species, Zootaxa 3913 (1), pp. 1-335 : 292

Galathea squamea Baba, 1979

Galathea squamea Baba, 1979a: 526 , figs 3, 4 (Noumea, New Caledonia, 20–23 m).—Baba et al., 2008: 77 (compilation).

Material examined. New Caledonia. Chesterfield Islands. CHALCAL 84, Stn D12, 20°31.33’S, 161°06.51’E, 80 m, 15 July 1984: 1 M 2.3 mm (MNHN-IU-2013-8254).—Stn CP12, 20°34.30’S, 158°47.40’E, 67 m, 23 July 1984: 9 M 2.8–3.3 mm, 1 ov. F 3.0– 3.1 mm, 7 F 2.4–3.0 mm (MNHN-IU-2013-8289).—Stn CP14, 21°13.50’S, 158°50.20’E, 66 m, 24 July 1984: 8 M 2.9–3.2 mm, 1 ov. F 3.0 mm, 6 F 2.7–3.1 mm (MNHN-IU-2013- 8255).—Stn CP15, 21°24.90’S, 159°09.30’E, 60 m, 25 July 1984: 4 M 3.6–3.8 mm (MNHN-IU-2013-8253).—Stn CP16, 21°41.67’S, 159°21.92’E, 53 m, 25 July 1984: 1 F 2.4 mm (MNHN-IU-2013-8264).—Stn D57, 21°29.50’S, 159°16.40'E, 62 m, 25 July 1984: 1 M 3.0 mm (MNHN-IU-2013-8270). CORAIL 2, Stn CP22, 20°33’S, 161°01’E, 85–88 m, 22 July 1988: 1 M 2.8 mm, 1 F 2.7 mm (MNHN-IU-2013-8267).—Stn DW35, 19°22’S, 158°53’E, 52 m, 23 July 1988: 1 M 2.4 mm (MNHN-IU-2013-8271). Senez Reef, 7 m, 7 September 1992: 2 M 2.7–2.8 mm, 3 ov. F 2.6–3.1 mm (MNHN-IU-2013-8265). Ouen Island, Stn 71, 22°20'S, 166°34'E, 22 m, August 1984: 1 ov. F 3.5 mm (MNHN-IU-2013-8282).—Stn 77, 22°25.9'S, 166°31.8'E, 22 m, August 1984: 3 M 3.0–4.0 mm, 2 ov. F 3.0– 3.8 mm (MNHN-IU-2013-8288).— Stn 94, 22°31'S, 166°33'E, 17 m, August 1984: 1 F 2.5 mm (MNHN-IU-2013- 8275).—Stn 123, 22°30'S, 166°40'E, 21 m, August 1984: 1 ov. F 3.1 mm (MNHN-IU-2013-8277).—Stn 127,22°31'S, 166°46'E, 55 m, August 1984: 2 M 3.0– 3.7 mm, 2 ov. F 3.4–3.5 mm (MNHN-IU-2013-8274).—Stn 161, 22°34'S, 166°38'E, 20 m, August 1984: 1 ov. F 3.2 mm (MNHN-IU-2013-8261).—Stn 243, 22°24'S, 167°01'E, 29 m, August 1984: 1 ov. F 3.2 mm (MNHN-IU-2013-8263).—Stn 248, 22°24'S, 166°47'E, 47 m, August 1984: 1 ov. F 3.3 mm (MNHN-IU-2013-8272). St. Vincent, Stn 164, 22°10'S, 166°09'E, 17 m, no date: 2 F 1.4–2.1 mm (MNHN-IU-2013-8266). Maitre Island, 25 m, 5 September 1978: 1 F 4.0 mm (MNHN-IU-2013- 8284).— 25 m, 19 September 1978: 1 M 2.6 mm, 1 ov. F 3.2 mm (MNHN-IU-2013-8252).— 20 m, June 1992: 4 M 3.1–3.6 mm, 4 ov. F 3.0– 3.4 mm (MNHN-IU-2013-8268).— 22°19.41'S, 166°20.89'E, 20 m, 9 November 1995: 3 M 1.4–1.6 mm, 2 F 1.4–1.5 mm (MNHN-IU-2013-8258).—Stn 82, 22°19.61'S, 166°24.07'E, 105 m, 14 November 1995: 4 M 1.6–2.5 mm, 6 ov. F 2.4–3.0 mm (MNHN-IU-2013-8262). South Reef, Stn 296, 22°41'S, 166°44'E, 26 m, November 1984: 1 ov. F 3.2 mm (MNHN-IU-2013-8285).—Stn 297, 22°38.9'S, 116°45.6'E, 30 m, November 1984: 2 M 2.8–2.9 mm (MNHN-IU-2013-8260).—Stn 304, 22°39.8'S, 166°47.9'E, 27 m, November 1984: 1 M 2.3 mm, 1 ov. F 2.8 mm (MNHN-IU-2013-8278). Noumea, 14 September 1978: 1 ov. F 3.1 mm (MNHN-IU-2013- 8251).—Stn 3, 22°21'S, 162°22'E, 15 m, 21 May 1984: 1 M 2.7 mm (MNHN-IU-2013-8279), 1 M 3.5 mm (MNHN-IU-2013-13993).—Stn 13, 22°19.6'S, 166°26.1'E, 20 m, 1 May 1984: 1 ov. F 3.1 mm (MNHN-IU-2013- 8283).—Stn 56, 22°10'S, 166°15'E, 11 m, May 1984: 3 ov. F 3.0– 3.8 mm, 1 F 2.8 mm (MNHN-IU-2013- 8280).—Stn 269, 22°18'S, 168°18'E, 20 m, November 1984: 2 ov. F 2.1–2.4 mm (MNHN-IU-2013- 8276).— 22°30'S, 166°26'E, 32 m, 8 November 1995: 1 M 2.4 mm, 1 ov. F 2.3 mm (MNHN-IU-2013-8257). Canards Island, 20 m, 29 March 1978: 2 ov. F 3.2–3.3 mm (MNHN-IU-2013-8287). Lagon, Stn 313, 22°40.3'S, 166°50.1'E, 30 m, November 1984: 2 M 1.8–2.6 mm (MNHN-IU-2013-8259).—Stn 336, 22°41.5'S, 166°51.4'E, 26 m, November 1984: 2 ov. F 3.1–3.2 mm (MNHN-IU-2013-8281).—Lagon, Goeland Island, 10 m, 16 April 1993: 1 F 2.2 mm (MNHN-IU-2013-8256). New Caledonia, 180 m, 30 June 1992: 1 M 2.1 mm, 2 F 2.4–3.0 mm (MNHN-IU-2013-8290-92). Croissant Reef, Stn 533, 20 m, 18 April 1994: 2 M 2.9–3.1 mm, 2 ov. F 2.7–2.8 mm (MNHN-IU-2013-8269). Plotmatre, 20 m, 10 November 1995: 1 M 2.3 mm, 1 ov. F 2.2 mm (MNHN-IU-2013- 8286). West Bank, no date: 3 M 1.6–2.3 mm, 3 ov. F 2.0–3.0 mm (MNHN-IU-2013-8273).

Remarks. The material examined agrees with the holotype male from New Caledonia (Baba 1990). However, we have observed a few specimens without epigastric spines (or very minute), as well as a few specimens with one minute parahepatic spine on each side. These specimens agree very well with the typical specimens in other diagnostic respects, and therefore, we regard that these differences represent intraspecific variation.

The genetic divergences of COI between G. squamea and all other species are usually larger than 15.0%. The smallest divergence of the same gene is observed in G. sanctae (6.3%) from New Caledonia and Vanuatu ( Tab. 3 )

Distribution. New Caledonia, 7– 180 m.