Cryptosomatula Bruce, 1940

Gimmel, Matthew L. & Leschen, Richard A. B., 2022, Revision of the genera of Picrotini (Coleoptera: Cryptophagidae: Cryptophaginae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1), pp. 61-109 : 80-82

Cryptosomatula Bruce, 1940

( Fig. 21 )

Cryptosomatula Bruce, 1940: 683 . Type species: Cryptosomatula longicornis Bruce, 1940 , by original monotypy.

Diagnosis. This genus may be distinguished by the antennal club comprised of three antennomeres, lack of tempora, evenly convex pronotum (without pits or impressions) with no teeth, platforms, or protruding anterior angles, the broad, flat prosternal process, and all five abdominal ventrites free. It appears to be closely related to Cryptothelypterus , from which it differs by the finely faceted eyes, long discrimen (longer than half length of metaventrite), narrow pronotum (narrower than elytra), and distinct elytral punctation.

Redescription. Length 2.20–2.43 mm. Body form ( Fig. 21 ) elongate, parallel-sided, shining dorsally, with moderately dense decumbent setae; unicolorous. Head without tempora; vertex without temporal depression immediately anterior to ridge; band of reticulate sculpture present. Frontoclypeus not projecting laterally; raised portion of frons between antennal insertions not constricted, wider than antennal club. Transverse ridge above antennal insertions absent. Eye large, rounded, finely faceted, contacting antennal cavity; interfacetal setae present. Antennal club consisting of 3 antennomeres; antenna inserted into small cavity; antennomere 9 subequal in width to antennomere 10. Mandible with apex bifid, subapical serrations present. Maxillary palpomere 4 distinctly longer than 3; palpomere 4 not subulate. Gena without antennal groove; genal spines subacute. Gular sutures incomplete, not reaching occipital foramen. Pronotum not explanate and not strongly constricted at base, narrower than elytra, widest at middle; anterior angles absent, not projecting anteriorly (not extending anterior to cervical foramen of prothorax), without a distinct flat glandular surface or platform; lateral carina present and complete, bearing weakly developed setigerous tubercles, without glabrous space, width of lateral bead wider than discal puncture and narrower than antennal funicle; disc with narrow, shallow transverse basal impression; paramedial carinae and paralateral plicae absent; posterolateral angles about right, not projecting. Prothoracic hypomeron separated from the prosternum by a suture. Prosternum with anterior margin on different plane as disc; prosternal process flattened, with lateral beads weak, process slightly expanded apically, broadly rounded and crenulate with minute setae; procoxal cavity with anterolateral notch. Scutellar shield clearly visible and transverse. Elytron without humeral tooth or impressions; subapical gape present; punctation confused, dense and moderately impressed; decumbent setae not forming undulate pattern, with postscutellar and subapical setae directed posteriorly. Hind wing well developed. Mesoventrite with mesoventrital cavity shallow and not flanked by sharp carinae. Mesanepisternal pit present and glabrous. Metaventrite without postcoxal lines; discrimen more than 1/2 length of metaventrite, posterior notch of metaventrite present. Metendosternite with anterior tendons approximate. Tarsi 5-5- 5 in female, 5-5- 4 in male; tarsi slender, tarsomere 5 as wide as preceding tarsomeres in lateral view; pro- and mesotarsomere 4 with ventral setae; mesotarsomere 3 unlobed, with few setae; mesotarsomeres 1–3 of equal lengths, mesotarsomere 4 slightly shorter, mesotarsomere 5 slightly shorter than mesotarsomeres 1–4 combined. Abdominal ventrites free and with medial and lateral calli present, intersegmental crenulations absent; ventrite 1 with intercoxal process broadly rounded, without postcoxal lines; medio-basal thickenings of ventrites 3–5 absent; apex of ventrite 5 without crenulations.Abdominal spiracles on segment VII with openings present and larger in diameter than spiracle VI, texture granulate and atrium rounded and saclike.

Remarks. This genus, like Cryptothelypterus , is known only from the Juan Fernandez Islands. Only three specimens are known. The male genitalia have not been described and are presumed lost (LൾඌർHൾඇ & Lൺඐඋൾඇർൾ 1991) and the specimens we studied (MNNC and NZAC) were females.

Biology. The single species has been collected at altitudes of 1000–1200 m by G. Kuschel.

Distribution. Chile (Juan Fernandez Islands).

Included species (1). Cryptosomatula longicornis Bruce, 1940 .

Figures

Figs 19–27. Habitus of species of Picrotini. 19 – Connatocryptus utiku sp. nov.; 20 – Cordosomatula magnabagishae sp. nov.; 21 – Cryptosomatula longicornis Bruce, 1940; 22 – Cryptothelypterus atomarioides (Grouvelle, 1919); 23 – Cryptothelypterus obrieni Leschen & Lawrence, 1991; 24 – Cryptothelypterus pteropilosus Leschen & Lawrence, 1991; 25 – Cryptothelypterus splendens (Bruce, 1940); 26 – Foveocryptus chenyandongi sp. nov.; 27 – Humerocryptus tumidus (Broun, 1893). Scale bars = 1 mm.