Euphthiracarus africanus, : Niedbala 1998 a, Niedbala & Liu, 2018

Niedbała, Wojciech & Liu, Dong, 2023, Systematic, synonymic and biogeographical list of ptyctimous mites (Acari, Oribatida) in the world (1799 – 2022), Zootaxa 5265 (1), pp. 1-442 : 18-29

Afrotropical, Tanzania, endemic africanus Hoplophthiracarus Mahunka, 1984 synonym of Plonaphacarus kugohi ( Aoki, 1959)

afromontanensis Oribotritia (Oribotritia) Niedbała, 2006

Oribotritia afromontanensis Niedbała, 2006

Oribotritia afromontanensis: Liu 2015 a, Niedbała & Liu 2018 .

Oribotritia (Oribotritia) afromontanensis: Subías 2022 .

South Africa: Western Cape, Mpumalanga ( Niedbała 2006c, Subías 2022).

South African, indigenous aggenitalis Euphthiracarus (Euphthiracarus) Aoki, 1984

Euphthiracarus aggenitalis Aoki, 1984

Euphthiracarus aggenitalis: Niedbała 2011b, 2012b , Niedbała & Liu 2018.

Euphthiracarus (Euphthiracarus) aggenitalis: Niedbała 2000 , Subías 2022.

Palaearctic: Japan ( Aoki 1984, Niedbała 2000, 2011b, 2012b, Subías 2022).

Eastern Palaearctic, Japan, endemic agulhasensis Notophthiracarus Niedbała, 2006

Notophthiracarus agulhasensis: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) agulhasensis: Subías 2022

South Africa: Western Cape ( Niedbała 2006c, Subías 2022).

South African, endemic aitu Plonaphacarus Liu et Zhang, 2015

Plonaphacarus aitu: Niedbała & Liu 2018 .

Phthiracarus (Archiphthiracarus) aitu: Subías 2022 .

Australasia: New Zealand ( Liu & Zhang 2015c, Subías 2017).

Australasian, New Zealand, endemic alajuela Oribotritia (Oribotritia) Niedbała, 2003

Oribotritia alajuela Niedbała, 2003

Oribotritia alajuela: Niedbała 2008e , Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) alajuela: Subías 2022 .

Neotropic: Costa Rica, Panama ( Niedbała 2003a, 2004a, 2008e, Subías 2022).

Neotropical, Mesoamerica alaskensis Maerkelotritia Hammer, 1967 synonym of Maerkelotritia kishidai (Aoki, 1958)

alazon Euphthiracarus (Euphthiracarus) Walker, 1965

Euphthiracarus alazon Walker, 1965

Euphthiracarus alazon: Niedbała 2002 a, Niedbała & Liu 2018 .

Euphthiracarus (Euphthiracarus) alazon: Subías 2022 .

Nearctic: USA (California) ( Marshall et al. 1987, Niedbała 2002a, Niedbała & Starý 2015e, Subías 2022, Walker 1965).

Nearctic, USA, endemic alienus Arphthicarus Niedbała, 2017

Arphthicarus alienus: Niedbała & Ermilov 2017b , Niedbała & Liu 2018.

Hoplophthiracarus (Notophthiracarus) alienus: Subías 2022 .

Neotropic: Antilles ( Trinidad) ( Niedbała & Ermilov 2017b, Subías 2022).

Neotropical, Trinidad, endemic alienus Notophthiracarus Niedbała, 1989

Notophthiracarus alienus: Niedbała 1992, 2000 , 2006d, Niedbała & Penttinen 2007, Liu & Zhang 2014 d, Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) alienus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) alienus: Subías 2022 .

Australasia: Australia (southern part) ( Balogh & Balogh 2002, Niedbała 1989b, 1992, 2000, 2006d, 2012a, 2017b, Niedbała & Colloff 1997, Niedbała & Penttinen 2007, Subías 2022).

Australasian, southern part of Australia alienus Phthiracarus Niedbała, 2022

Phthiracarus alienus Niedbała, 2022 : Niedbała & Ermilov 2022b.

Neotropic: Colombia ( Niedbała & Ermilov 2022b).

Neotropical, Colombia, endemic aliquantus Phthiracarus Niedbała, 1988

Phthiracarus (Archiphthiracarus) aliquantus: Subías 2022 .

Phthiracarus aliquantus: Niedbała 1992, 2002a , 2007c, Niedbała & Liu 2018.

Nearctic: Canada (Yukon, British Columbia), USA (Oregon, Washington, Arizona, Kentucky, California, Tennessee, Pennsylvania) ( Niedbała 1988e, 1992, 2002a, 2007c, Niedbała & Starý 2015e, Subías 2022). Nearctic, western part with eastern disjunction (Kentucky)

alius Arphthicarus Niedbała, 2019

Arphthicarus alius: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) alius: Subías 2022 .

Afrotropic: Tanzania ( Zanzibar) ( Niedbała & Ermilov 2019b).

Afrotropical, Tanzania, endemic alius Steganacarus (Rhacaplacarus) Niedbała, 2018

Hoplophorella (Rhacaplacarus) alius: Subías 2022 .

Afrotropic: Cameroon ( Niedbała & Osiejuk 2018).

Afrotropical, Cameroon, endemic allocota Oribotritia (Oribotritia) Niedbała, 2003

Oribotritia allocota Niedbała, 2003

Oribotritia allocota: Niedbała 2004a, 2008e , Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) allocota: Subías 2022 .

Neotropic: Costa Rica, Colombia, Venezuela ( Niedbała 2003a, 2004a, 2008e, Subías 2022).

Neotropical, Mesoamerica and northern part of South America allocotos Arphthicarus Niedbała, 2003

Arphthicarus allocotos: Niedbała 2004a, 2008e , Liu & Oconnor 2014, Niedbała & Liu 2018. Hoplophthiracarus (Notophthiracarus) allocotos: Subías 2022 .

Neotropic: Costa Rica, Panama, Venezuela ( Niedbała 2003a, 2004a, 2008e, Subías 2022).

Neotropical, Mesoamerican and Guyano-Brasilian allocotos Indotritia (Indotritia) Niedbała, 2004

Indotritia allocotos Niedbała, 2004

Indotritia allocotos: Niedbała & Liu 2018 .

Indotritia (Indotritia) allocotos: Liu 2015c , Oliveira et al. 2017, Subías 2022.

Neotropic: Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Brazil, endemic allocotos Phthiracarus Niedbała et Starý, 2015

Phthiracarus allocotos: Niedbała & Liu 2018 .

Phthiracarus (Neophthiracarus) allocotos: Subías 2022 .

Neotropic: Bolivia ( Niedbała & Starý 2015b, Subías 2022).

Neotropical, Brazil, endemic alpinus Notophthiracarus Balogh et Mahunka, 1997 synonym of Notophthiracarus sinuosus (Niedbała, 1982) altissima Microtritia Mahunka, 1994 synonym of Microtritia hauseri Mahunka, 1993

altitudinis Steganacarus (Tropacarus) Bernini, Bernini et Avanzati, 1989 synonym of Steganacarus (Tropacarus) desmeti Niedbała, 1986

amatholensis Phthiracarus Niedbała, 2021

Phthiracarus (Archiphthiracarus) amatholensis: Subías 2022 .

South Africa: Eastern Cape (Niedbała et al. 2021).

South African, endemic amazonensis Mesotritia ( Märkel, 1964)

Perutritia amazonensis Märkel, 1964

Mesotritia amazonensis: Niedbała 2004a , Liu & Chen 2010 a, Niedbała & Liu 2018.

Mesotritia (Perutritia) amazonensis: Subías 2022 .

Neotropic: Peru ( Balogh & Balogh 1987, 1988, 2002, Märkel 1964, Niedbała 2004a, Subías 2022). Neotropical, Peru, endemic ambiguus Hoplophthiracarus Niedbała, 1982

Hoplophthiracarus ambiguus: Niedbała 1986b, 1992 , 2001 a, Niedbała & Liu 2018.

Rhacaplacarus (Rhacaplacarus) ambiguus: Subías 2004 .

Hoplophorella (Rhacaplacarus) ambigua: Subías 2022 .

Afrotropic: Gabon, Democratic Republic Congo, Ethiopia, Uganda, Kenya, Tanzania ( Balogh & Balogh 2002, Niedbała, 1982f, 1986b, 1992, 2001a, 2008c, 2017c, Niedbała & Ermilov 2011, 2012, 2014a, 2022c, Niedbała & Starý 2015e, Subías 2022).

Afrotropical, central part, continuous amboroensis Protophthiracarus Niedbała et Starý, 2015

Protophthiracarus amboroensis: Niedbała & Liu 2018 .

Phthiracarus (Neophthiracarus) amboroensis: Subías 2022 .

Neotropic: Bolivia ( Niedbała & Starý 2015b, Subías 2022).

Neotropical, Bolivia, endemic americanus Phthiracarus Ewing, 1909 synonym of Acrotritia ardua (C.L. Koch, 1841)

amoenus Steganacarus (Rhacaplacarus) (Niedbała, 1983)

Hoplophorella amoena Niedbała, 1983

Steganacarus (Rhacaplacarus) amoenus: Niedbała 1986b, 1992 , 2001a, Subías 2004, Niedbała & Liu 2018. Hoplophorella (Rhacaplacarus) amoena: Subías 2022 .

Afrotropic: Uganda ( Balogh & Balogh 2002, Niedbała 1983h, 1986b, 1992, 2001a, Subías 2022). Afrotropical, Uganda, endemic ampla Oribotritia (Oribotritia) Niedbała, 1991

Oribotritia ampla Niedbała, 1991

Oribotritia ampla: Niedbała 1998b , Liu 2015a, 2016b, Subías 2004, Niedbała & Liu 2018.

Oribotritia (Oribotritia) ampla: Subías 2022 .

Australasia: Fiji, Samoa Islands ( Balogh & Balogh 2002, Niedbała 1991d, 1998b, 2012a, Subías 2022). Australasian, Pacific Islands ampliata Oribotritia (Oribotritia) Niedbała, 2004

Oribotritia ampliata Niedbała, 2004

Oribotritia ampliata: Liu 2015 a, Niedbała & Liu 2018 .

Oribotritia (Oribotritia) ampliata: Subías 2022 .

Neotropic: Jamaica ( Niedbała 2004a, Subías 2022).

Neotropical, Jamaica, endemic amus Austrophthiracarus Niedbała et Starý, 2014

Austrophthiracarus amus: Liu & Zhang 2016 a, Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) amus: Subías 2022

Neotropic: Costa Rica ( Niedbała & Starý 2014e, Subías 2022).

Neotropical, Costa Rica, endemic anakolos Phthiracarus Niedbała, 2006

Phthiracarus anakolos: Niedbała & Liu 2018 .

Hoplophthiracarus (Steganacarellus) anakolos: Subías 2022 .

South Africa: Eastern Cape ( Niedbała 2006c, Subías 2022).

South African, endemic anceps Austrophthiracarus Niedbała, 2004

Austrophthiracarus anceps: Niedbała 2008e , Liu & Zhang 2016 a, Niedbała & Liu 2018. Hoplophthiracarus (Notophthiracarus) anceps: Subías 2022 .

Neotropic: Antilles ( Cuba) ( Niedbała 2004a, 2008e, Subías 2022).

Neotropical, Cuba, endemic anceps Oribotritia (Oribotritia) Niedbała, 2000

Oribotritia anceps Niedbała, 2000

Oribotritia anceps: Subías 2004 , Niedbała 2011b, 2012b, Liu 2015a, Subías 2004, Niedbała & Liu 2018, Liu 2021b.

Oribotritia (Oribotritia) anceps: Subías 2022 .

Orient: China (Jiangxi). Palaearctic: Nepal (Liu 2021b, Liu et al. 2011a, Niedbała 2000, 2011b, 2012b, Niedbała & Ermilov 2018b, Subías 2022).

Oriental (Chinese), introduced to Nepal anchistea Rhysotritia Niedbała, 1998 synonym of Acrotritia vestita ( Berlese, 1913)

andamanensis Arphthicarus Niedbała, 2011

Arphthicarus andamanensis: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) andamanensis: Subías 2022 .

Orient: Andaman Islands ( Niedbała 2011a, Subías 2022).

Oriental, Andaman Islands, endemic andasibensis Notophthiracarus Niedbała et Starý, 2014

Notophthiracarus andasibensis: Niedbała & Liu 2018 .

Phthiracarus (Archiphthiracarus) andasibensis: Subías 2022 .

Afrotropic: Madagascar (Niedbała 2017 a, Niedbała & Starý 2014b, Subías 2022).

Afrotropical, Madagascar, endemic andinus Austrophthiracarus ( Balogh, 1984)

Phthirarica andina Balogh, 1984

Phthirarica andina: Niedbała 1992 .

Notophthiracarus (Protophthiracarus) andinus: Subías 2004 .

Hoplophthiracarus (Protophthiracarus) andinus: Subías 2022 .

Austrophthiracarus andinus: Niedbała 1994a, 2004a , Liu & Zhang 2016 a, Niedbała & Liu 2018. Neotropic: Antilles ( Cuba), Colombia ( Balogh 1984, Balogh & Balogh 1987, 1988, 2002, Niedbała 1992, 1994a, 2004a, Niedbała & Ermilov 2015, Subías 2022).

Neotropical, Mesoamerica andreasi Phthiracarus Niedbała, 2020

Palaearctic: Azores ( Niedbała & Buczkowska 2020).

Palaearctic, Azores, endemic andrei Atropacarus (Hoplophorella) ( Balogh, 1958)

Steganacarus andrei Balogh, 1958

Steganacarus andrei: Niedbała 1986b, 1992 .

Hoplophorella andrei: Mahunka 1984b , Pérez-Íñigo & Baggio 1993, Niedbała & Schatz 1996, Niedbała & Corpuz-Raros 1998, Niedbała 2001a, b.

Hoplophorella (Hoplophorella) andrei ( Balogh, 1958) : Subías 2022.

Atropacarus (Hoplophorella) andrei: Niedbała 1998b , Niedbała & Corpuz-Raros 1998, Liu & Oconnor 2015, Niedbała & Liu 2018, Niedbała & Ermilov 2022b.

Atropacarus (Hoplophorella) vitrinus (in part): Niedbała 2000.

Atropacarus (Hoplophorella) nigeriensis Badejo, 2001 : in Badejo, Woas & Beck 2001a: Niedbała & Liu 2018.

Palaearctic: Germany (Berlin, Palm house), Poland (Palm House), Czech Republik, Praha, Palm house). Afrotropic: Ivory Coast, Cameroon, Nigeria (?), Angola, Malawi, Seychelles. Orient: Philippines. Neotropic: Galapagos Islands, Ecuador, Colombia, Peru ( Badejo et al. 2001a, Balogh 1958, Ermilov et al. 2016, Liu & Oconnor 2015, Mahunka 1984b, Niedbała 1986b, 1998b, 2001a, 2008c, 2014c, 2015b, 2017a, 2018, Niedbała & Ermilov 2017a, 2019a, 2022b, Niedbała & Osiejuk 2018, Niedbała & Roszkowska 2017, Niedbała & Starý 2022, Schatz 1998, Subías 2022).

Pantropical, only in Afrotropical and Neotropical Regions, introduced to Palm houses in Berlin, Praha and in Poland

Note. Subías (2022) incorrectly synonymized A. (H.) andrei with Hoplophorella lienhardi Mahunka, 1987 which should be synonym of A. (H.) vitrinus .

angolensis Atropacarus (Hoplophorella) (Mahunka, 1984)

Hoplophorella angolensis Mahunka, 1984

Hoplophorella angolensis: Niedbała 1992 .

Rhacaplacarus angolensis: Mahunka 1992 .

Atropacarus (Hoplophorella) angolensis: Niedbała 2001 a, Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) angolensis: Subías 2022 .

Afrotropic: Angola ( Balogh & Balogh 2002, Mahunka 1984 b, 1985, 1992, Niedbała 1992, 2001a, Subías 2022).

Afrotropical, Angola, endemic angolensis Phthiracarus Mahunka, 1985 synonym of Phthiracarus lentulus (C.L. Koch, 1841)

angulatus Steganacarus (Steganacarus) Balogh et Mahunka, 1992

Neosteganacarus angulatus Balogh et Mahunka, 1992

Steganacarus (Steganacarus) angulatus: Niedbała 2004 a, Niedbała & Liu 2018 .

Steganacarus (Rafacarus) angulatus: Subías 2004 .

Atropacarus (Rafacarus) angulatus: Subías 2022 .

Neotropic: Brazil ( Balogh & Balogh 2002, Balogh & Mahunka 1992, Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Brazil, endemic angusta Oribotritia (Oribotritia) Mahunka, 1982

Oribotritia angusta Mahunka, 1982

Oribotritia angusta: Niedbała 2000, 2011b , 2012b, Subías 2004, Liu et al. 2011a, Liu 2015a, 2021b, Niedbała & Liu 2018.

Oribotritia (Oribotritia) angusta: Subías 2022 .

Palaearctic: North Korea, China (Beijing, Gansu, Hebei, Shanxi, Inner Mongolia), Russia (Far East) ( Chen et al. 2013, Liu 2015a, 2021b, Liu et al. 2011a, Mahunka 1982c, Niedbała 2000, 2011b, 2012b, 2015b, Subías 2022).

Eastern Palaearctic, known from Amurian and Manchurian areas and from Northern China region angustatus Hoplophthiracarus Niedbała, 1984

Hoplophthiracarus angustatus: Niedbała 1986b, 1992 , 2000, 2011b, 2012b, 2018a, Subías 2004, Niedbała & Liu 2018.

Hoplophthiracarus (Hoplophthiracarus) angustatus: Subías 2022 .

Palaearctic: Nepal ( Balogh & Balogh 2002, Niedbała 1984d, 1986b, 1992, 2000, 2011b, 2012b, Niedbała & Ermilov 2018a, Subías 2022).

Southern Palaearctic, Nepal, endemic angustus Notophthiracarus Niedbała, 2012

Notophthiracarus angustus: Liu & Zhang 2014 d, Niedbała & Liu 2018 .

Hoplophthiracarus (Besuchetacarus) angustus: Subías 2022 .

Australasia: Australia (New South Wales) ( Niedbała 2012a, Subías 2022).

Australasian, Australia, endemic anomala Hoplophora Berlese, 1883 synonym of Steganacarus (Steganacarus) magnus ( Nicolet, 1855)

Note. Subías (2022) considered this species as subspecies of S. (S.) magnus although it was previously synonymized with S. (S.) magnus in Bernini & Avanzati (1988a).

anonymus Phthiracarus Grandjean, 1933

Phthiracarus (Archiphthiracarus) anonymus: Subías 2022 .

Phthiracarus anonymum: Grandjean 1933a , d, 1934a, 1939b, 1940b, 1941c, 1942a, 1946a, b, 1949, 1950, 1954a, 1958, 1964a, b, 1965, 1967.

Phthiracarus anonymus: Hammen 1965 , Krivolutsky 1975, Parry 1979a, Pérez-Íñigo 1979, Niedbała et al. 1981, 1982, 1990, Balogh & Mahunka 1983, Niedbała 1986b, 1992, 1994c, 2000, 2001a, 2002a, b, 2003a, 2004a, d, 2006c, d, 2007a, 2008e, f, 2010b, 2011b, 2012b, 2015b, Marshall et al. 1987, Berg et al. 1990, Niedbała & Schatz 1996, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Mahunka & Mahunka-Papp 2004, Weigmann 2006, Niedbała & Illig 2007b, Liu & Zhang 2016b, Niedbała & Liu 2018, Niedbała & Starý 2022.

Archiphthiracarus anonymus: Balogh & Mahunka 1980 a, 1983 .

Phthiracarus (Archiphthiracarus) anonymus: Subías 2004 .

Phthiracarus anonymus amicus: Niedbała 1986b, 1992 .

Phthiracarus anonymus amicus Jacot, 1938 syn. by Niedbała 2002a.

Archiphthiracarus hauseri: Niedbała 1992 .

Archiphthiracarus hauseri Mahunka, 1988 syn. by Niedbała 2001a.

non Phthiracarus anonymus sensu Feider & Suciu, 1957

Nearctic: USA (New York, Arizona, California, North Carolina). Bermuda. Palaearctic: Azores, Madeira, Scotland, England, Portugal, Spain, France, Belgium, the Netherlands, Denmark, Germany, Luxembourg, Austria, Italy, Poland, Hungary, Greece, Georgia, Algeria, Turkey. Neotropic: Jamaica, Mexico, Guatemala, Costa Rica, Panama, Galapagos Islands, Ecuador, Venezuela, Bolivia, Peru, Brazil. Afrotropic: Cameroon, Ethiopia, Uganda, Tanzania, Malawi, Madagascar, Reunion. South Africa: Mpumalanga, Free State, KwaZulu-Natal, Western Cape. Orient: Sri Lanka, Java, Malaysia ( Sarawak). Australasia: Australia (southern part), New Zealand, Hawaii Islands ( Balogh & Balogh 2002, Balogh & Mahunka 1980 a, 1983, Beck et al. 2014, Berg et al. 1990, Grandjean 1933a, d, 1934a, 1939b, 1940b, 1941c, 1942a, 1946a, b, 1949, 1950, 1954a, 1958, 1964a, b, 1965, 1967, Hammen 1965, Jacot 1938, Mahunka 1983a, 1988b, 1993d, 2002, Mahunka & Paoletti 1984, Mahunka et al. 2013, Marshall et al. 1987, Morell & Subías 1991, Niedbała 1986b, 1992, 1994c, 1998c, 2000, 2001a, 2002b, 2002a, 2003a, 2004a, 2006c, d, 2007a, 2008b, e, f, 2010b, 2012a, 2011b, 2012b, 2015b, 2017a, c, 2018, Niedbała & Buczkowska 2020, Niedbała & Dabert 2013, Niedbała & Ermilov 2011, 2016, 2022c, Niedbała et al. 2021, Niedbała & Illig 2007b, 2018b, Niedbała & Roszkowska 2017, Niedbała & Schatz 1996, Niedbała & Starý 2014d, 2015b, e, 2016a, 2022, Norton & MacNamara 1976, Oliveira 2004, Oliveira et al. 2017, Olszanowski et al. 1996, Parry 1979a, Pérez-Íñigo 1979, Schatz 1983, 1998, Seniczak et al. 2021, Subías 2022, Weigmann et al. 2015).

Semicosmopolitan, Panpalaearctic, except central and eastern asiatic provinces anonymus amicus Phthiracarus Jacot, 1938 = Phthiracarus anonymus Grandjean, 1933

anosculpturatus Austrophthiracarus (Mahunka, 1987)

Hoplophthiracarus anosculpturatus Mahunka, 1987

Hoplophthiracarus anosculpturatus: Niedbała 1992 .

Calyptophthiracarus anosculpturatus: Mahunka 1992 .

Austrophthiracarus anosculpturatus: Niedbała 2001a , Liu & Zhang 2016 a, Niedbała & Liu 2018.

Phthiracarus (Neophthiracarus) anosculpturatus: Subías 2004 .

Hoplophorella (Hoplophorella) anosculpturata: Subías 2022 .

Afrotropic: Kenya ( Balogh & Balogh 2002, Mahunka 1987b, 1992, Niedbała 1992, 2001a, Subías 2022).

Afrotropical, Kenya, endemic antennatus Steganacarus Mahunka et Mahunka-Papp, 2003 synonym of Steganacarus (Steganacarus) vernaculus Niedbała, 1982

antipai Euphthiracarus (Euphthiracarus) Feider et Călugăr, 1968

Euphthiracarus antipai Feider et Călugăr, 1968

Euphthiracarus antipai: Balogh & Mahunka 1980 a, 1983 , Niedbała 2011b,

2012b, Niedbała & Liu 2018.

Euphthiracarus (Euphthiracarus) antipai: Subías 2022 .

Palaearctic: Romania ( Balogh & Mahunka 1980 a, 1983, Feider & Călugăr 1968, Niedbała 2011b, 2012b, Subías 2022).

Western Palaearctic, Romania, endemic

Note. This species differs from Euphthiracarus (Euphthiracarus) monodactylus ( Willmann, 1919) only in the absence of exobothridial setae ( Mahunka & Mahunka-Papp 1995, p. 38).

antrosus Atropacarus (Atropacarus) Niedbała, 2003

Atropacarus (Atropacarus) antrosus: Niedbała 2004a, 2008e , Niedbała & Liu 2018, Subías 2022.

Neotropic: Costa Rica, Panama ( Niedbała 2003a, 2004a, 2008e, Subías 2022).

Neotropical, Mesoamerica aokii Acrotritia ( Niedbała, 2000)

Rhysotritia aokii Niedbała, 2000

Acrotritia aokii: Subías 2004 , Niedbała 2011b, 2012b, Liu 2015d, 2021b, Niedbała & Liu 2018.

Afrotropic: Kenya. Palaearctic: Turkey, India (Himalah Pradesh), Nepal, Russia (Far East), China (Tibet), North and South Korea, Japan. Orient: Northern Vietnam, Indonesia (Sulawesi, Sumatra) ( Ermilov et al. 2012, Kaczmarek & Niedbała 2002, Liu 2021b, Niedbała 2000, 2008c, 2011b, 2012b, 2015b, Niedbała & Ermilov 2013 a, 2014b, Subías 2004, Vu 2013).

Semicosmopolitan, except Nearctic and Neotropical Regions; in Palaearctic Region, only in southern and eastern parts of Asiatic provinces

Note. Subías (2022) considered this species as congeneric with A. sinensis Jacot, 1923 which is highly probable.

aokii Apoplophora Mondal, Kundu et Roy, 1999

Apoplophora aokii: Liu et al. 2009a , Liu 2016 a, Niedbała & Liu 2018, Subías 2022.

Orient: India (West Bengal, Darjeeling) ( Mondal et al. 1999, Subías 2022).

Oriental, India, endemic aokii Arphthicarus ( Niedbała, 1987)

Austrophthiracarus aokii Niedbała, 1987

Austrophthiracarus aokii: Niedbała 1992 .

Arphthicarus aokii: Niedbała 1994a, 2000 , Liu & Zhang 2015c, Niedbała & Liu 2018.

Phthiracarus (Neophthiracarus) aokii: Subías 2004 .

Hoplophthiracarus (Protophthiracarus) aokii: Subías 2022 .

Australasia: Australia (Queensland) ( Balogh & Balogh 2002, Niedbała 1987, 1992, 2000, 2012a, Subías 2022).

Australasian, Australia, endemic aokii Austrophthiracarus (Mahunka, 1983)

Hoplophthiracarus aokii Mahunka, 1983

Hoplophthiracarus aokii: Niedbała 1986b, 1992 .

Calyptophthiracarus aokii: Mahunka 1992 .

Austrophthiracarus aokii: Niedbała 2001a , Liu & Zhang 2016 a, Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) aokii: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) aokii: Subías 2022 .

Afrotropic: Tanzania, Malawi ( Balogh & Balogh 2002, Mahunka 1983a, 1986, 1992, Niedbała 1986b, 1992, 2001a, 2017c, Subías 2022).

Afrotropical, eastern part

Note. Subías (2022) considered Austrophthiracarus aokiii malagasensis Mahunka, 1983 as subspecies of this species. See note of Notophthiracarus malagasensis (Mahunka, 2010) .

aokii Oribotritia (Oribotritia) Mahunka, 1987

Oribotritia aokii Mahunka, 1987

Oribotritia aokii: Niedbała & Corpuz-Raros 1998 , Niedbała 2000, Subías 2004, Niedbała et al. 2006, Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) aokii: Subías 2022 .

Orient: Malaysia (Sabah, Pahang), Indonesia, Philippines ( Balogh & Balogh 2002, Mahunka 1987 d, Niedbała 2000, Niedbała & Corpuz-Raros 1998, Niedbała & Ermilov 2017 a, Niedbała et al. 2006, Niedbała & Starý 2015e, Subías 2022).

Oriental, Sundaian and Philippinean aotearoana Indotritia (Zeaotritia) Ramsay, 1966

Indotritia aotearoana Ramsay, 1966

Indotritia aotearoana: Niedbała 2006d .

Indotritia (Zeaotritia) aotearoana: Liu 2015c , Niedbała & Liu 2018, Subías 2022.

Australasia: New Zealand, Australia (southern part) ( Balogh & Balogh 2002, Niedbała 2000, 2006d, 2012a, Ramsay 1966, Subías 2022).

Australasian, New Zealand and southern part of Australia apiculatus Phthiracarus Jacot, 1939 synonym of Phthiracarus longulus (C.L. Koch, 1841)

applicatus Steganacarus (Steganacarus) ( Sellnick, 1920)

Phthiracarus applicatus Sellnick, 1920

Hoploderma applicata (- um): Sellnick 1928, Willmann 1931.

Phthiracarus applicatus: Sellnick 1922 , Castagnoli & Pegazzano 1985.

Hoplopderma applicata (- um): Sellnick 1928, Willmann 1931.

Steganacarus applicatus: Jacot 1936 , Hammen 1959, Sellnick 1960, Krivolutsky 1975, Balogh & Mahunka

1980 a, 1983, Bernini & Avanzati 1989 c, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008.

Steganacarus (Steganacarus) applicatus: Doumandji et al. 1985 , Niedbała 1985b, 1986b, 1992, 2008f, 2015b, Avanzati et al. 1995, Subías 2022, 2011b, 2012b, Weigmann 2006, Niedbała & Liu 2018, Niedbała et al. 2021.

Phthiracarus berlesei Oudemans, 1915 syn. by Subías 2004 (?).

Hoplophora quadriseriata Haupt, 1882 syn. by Subías 2004 (?).

Palaearctic: Madeira, Portugal, Spain, Algeria, Norway, Sweden, Finland, France, Germany, Switzerland, Italy, Austria, Poland, Czech Republik, Croatia, Slovenia, Estonia, Russia (Central European Territory) ( Avanzati et al. 1995, Balogh & Mahunka 1980 a, 1983, Bernini &Avanzati 1989c, Castagnoli & Pegazzano 1985, Doumandji et al. 1985, Hammen 1959, Jacot 1936, Mahunka et al. 2013, Mahunka & Mahunka-Papp 2003, Niedbała 1985b, 1986b, 1992, 2008f, 2011b, 2012b, 2015b, Niedbała et al. 2020, 2021, Olszanowski et al. 1996, Schatz 1983, Sellnick 1920, Seniczak et al. 2021, Subías 2022, Weigmann et al. 2015).

Western Palaearctic, Paneuropean and also reported from the Maghrebian region

Note. It is not understandable why Subías (2004) synonymized Phthiracarus berlesei Oudemans, 1915 and Hoplophora quadriseriata Haupt, 1882 with S. (S.) applicatus .

aquilus Notophthiracarus Niedbała, 2000

Notophthiracarus aquilus: Liu & Zhang 2013d, 2015b , Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) aquilus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) aquilus: Subías 2022 .

Australasia: New Zealand ( Niedbała 2000, 2012a, Subías 2022).

Australasian, New Zealand, endemic araios Protophthiracarus Niedbała, 2001

Protophthiracarus araios: Niedbała 2004 d, Niedbała & Liu 2018 .

Notophthiracarus (Notophthiracarus) araios: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) araios: Subías 2022 .

Afrotropic: Madagascar ( Mahunka 2002, Niedbała 2001a, 2004d, 2017a, Niedbała & Starý 2016a, Subías 2022).

Afrotropical, Madagascar, endemic arctata Hoplophora Riley, 1874 synonym of Acrotritia ardua (C.L. Koch, 1841)

ardua Acrotritia (C.L. Koch, 1841)

Hoplophora ardua C.L. Koch, 1841 .

Hoplophora ardua: Grandjean 1953 .

Pseudotritia ardua: Grandjean 1933a , d, 1934b, 1935, 1939a, b, 1940a, b, 1941a, b, 1942a, b, 1949, 1954a, 1958, Hammen 1952, 1959, Aoki 1958b, 1968b.

Rhysotritia ardua ardua: Pérez-Íñigo 1969a, 1974 , Balogh & Mahunka 1983, Mahunka 1990b, Mahunka & Mahunka-Papp 2004.

Rhysotritia ardua: Grandjean 1964a , b, 1965, 1966, 1967, 1968, 1969, Märkel 1964, Pérez-Íñigo 1969c, Krivolutsky 1975, Balogh & Mahunka 1980 a, 1983, Niedbała et al. 1981, 1982, 1990, Mahunka 1982c, Doumandji et al. 1985, Niedbała 1985b, 1993a, 1994c,1995b, 1998a, b, 2000, 2001a, b, 2002a, b, 2003a, 2004a, c, d, Marshall et al. 1987, Niedbała & Schatz 1996, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004a, 2008, Shimano & Aoki 1997a, Niedbała & Corpuz-Raros 1998, Baran & Ayyildiz 2000, Olszanowski & Niedbała 2000, Wang et al. 2002, Weigmann 2006, Mortazavi et al. 2010.

Acrotritia ardua: Niedbała 2006c , d, 2007a, c, 2008e, f, 2010b, 2011a, b, 2012b, Niedbała & Penttinen 2007, Chen et al. 2010, Liu & Zhang 2014a, Liu 2015d, 2021b, Liu & Chen 2015, Niedbała & SzywilewskaSzczykutowicz 2017, Niedbała & Ermilov 2018a, 2020, 2022b, Niedbała & Liu 2018, Ryabinin et al. 2018, Niedbała et al. 2021, Subías 2022.

Phthiracarus arduus: Castagnoli & Pegazzano 1985 .

Acrotritia ardua ardua: Subías 2004 .

Pseudotritia ardua antetriheterodectylum: Jacot 1939 .

Pseudotritia ardua obesum: Jacot 1930 .

Pseudotritia ardua monoheterodactylum: Jacot 1930 .

Pseudotritia ardua triheterodactylum: Jacot 1930 .

Rhysotritia ardua penicillata Pérez-Íñigo, 1969 syn. by Mahunka & Mahunka-Papp 1995, 2004.

Rhysotritia ardua penicillata: Balogh & Mahunka 1980 a, 1983 , Mahunka & Mahunka-Papp 2004.

Phthiracarus americanus Ewing, 1909 syn. by Jacot 1938.

Phthiracarus americanus: Castagnoli & Pegazzano 1985 .

Acrotritia ardua americana: Subías 2022 .

Hoplophora arctata Riley, 1874 syn. by Oudemans 1916a.

Rhysotritia arctata: Mahunka & Mahunka-Papp 2004 .

Phthiracarus canestrini Michael, 1898 syn. by Oudemans 1900.

Phthiracarus canestrini: Hammen 1959 .

Tritia canestrini: Castagnoli & Pegazzano 1985 .

Tritia lentula Canestrini, 1885 syn. by Oudemans 1916a.

Tritia lentula (K.) var. hyeroglyphica Berlese, 1916 syn. by Niedbała 1993a.

Tritia lentula (K.) var. hyeroglyphica: Castagnoli & Pegazzano 1985 .

Rhysotritia ardua hyeroglyphica: Mahunka 1991 a, Mahunka & Mahunka-Papp 1995, 2003 , 2004.

Phthiracarus pectinatus Ewing, 1917 syn. by Jacot 1930.

Pseudotritia loricata ( Rathke, 1799) syn. by Feider & Suciu 1957.

Trombidium loricatum: Subías 2017 .

Rhysotritia ardua affinis Sergienko, 1989 syn. by Niedbała 2011b.

Acrotritia ardua affinis: Subías 2004 .

Rhysotritia hauseri Mahunka, 1991 syn. by Subías 2004.

Rhysotritia pinguisetosa Bayartogtokh, 2010 syn. by Niedbała 2015b.

Rhysotritia pinguisetosa Bayartogtokh, 2010 : Subías 2022.

Acrotritia pinguisetosa: Liu 2015d .

Nearctic: Canada (North-West Territories, Yukon, Alberta, Saskatchewan, Manitoba, Ontario, New Brunswick, Nova Scotia , Quebec), USA (Illinois, Wyoming, Colorado, Arizona, Montana, South Dakota, Nebraska, Oklahoma, Texas, Minnesota, Wisconsin, Iowa, Missouri, Illinois, Michigan, Indiana, Ohio, Kentucky, Maine, Vermont, New Hampshire, Massachussetts, Pennsylvania, New York, New Jersey, West Virginia, Arkansas, Louisiana, Mississippi, Tennessee, Alabama, North Carolina, Maryland, Virginia, South Carolina, Georgia, Florida), Bermuda. Palaearctic: Azores, Portugal, Spain, France, Denmark, Norway, Sweden, Finland, the Netherlands, Germany, Luxembourg, Switzerland, Italy, Austria, Poland, Czech Republic, Slovakia, Hungary, Croatia, Bosnia-Herzegovina, Macedonia, Romania, Greece, Turkey, Estonia, Latvia, Lithuania, Belarus, Ukraine (western part and Crimea), Azerbaijan, Georgia, Russia (North, Central and South European Territories, Western Siberia), Mongolia, Canary Islands, Morocco, Algeria, Turkey, Syria, Iraq, Iran, Kazakhstan, Uzbekistan, Turkmenistan, Kyrgyzstan, India (Arunachal Pradesh, northern part), Nepal, Russia (Far East), North and South Korea, China (Jilin, Heilongjiang), Japan. Neotropic: Cocos Island, Mexico, Guatemala, Belize, Costa Rica, Panama, Antilles, Galapagos Islands, Colombia, Venezuela, Bolivia, Brazil, Chile, Uruguay. Afrotropic: Ivory Coast, Ghana, Togo, Cameroon, Democratic Republic Congo, Ethiopia, Kenya, Somalia, Rwanda, Tanzania, Angola, Zambia, Malawi, Comoro Islands, Madagascar (Nguru Mts., Zanzibar). South Africa: Limpopo, North West, Gauteng, Mpumalanga, Free State, KwaZulu-Natal, Western and Eastern Cape. Orient: North and South India, Nepal, Sri Lanka, Southern and Northern Vietnam, Cambodia, China (Taiwan), Malaysia (Sabah, West Malaysia), Indonesia (Sumatra, Sumbawa), Philippines. Australasia: Papua New Guinea, Australia (northern and southern parts), New Caledonia, Tonga, Samoa Islands, Society Islands, Marquesas Islands, Henderson Island, Easter Island, Hawaii ( Ahaniazad et al 2016, Aoki 1958b, 1970, 1980, Ayyildiz & Özkan 1988, Ayyildiz et al. 2011, Balogh & Balogh 2002, Balogh & Mahunka 1980 a, 1983, 2002, Baran & Ayyildiz 2000, Bayartogtokh 2010, Castagnoli & Pegazzano 1985, Chen et al. 2010, 2013, Doumandji et al. 1985, Ermilov 2011, Ermilov & Niedbała 2013, 2014 a, Ermilov et al. 2012, Ewing 1909a, 1917a, Feider & Suciu 1957, Grandjean 1933a, d, 1934b, 1935, 1939a, b, 1940a, b, 1941a, b, 1942a, b, 1949, 1953, 1954a, 1958, 1966, 1967, 1968, 1969, Hammen 1952, 1959, Hammer 1952, Kaczmarek & Niedbała 2002, Lions 1964, 1967, Liu 2015d, 2021b, Liu & Chen 2015, Mahunka 1982c, 1990b, 1991a, 1993, Mahunka et al. 2013, Mahunka & Mahunka-Papp 1995, 1999, 2003, 2004, Mahunka & Paoletti 1984, Märkel 1964, Marshall et al. 1987, Mortazavi et al. 2010, Niedbała 1985b, 1993a, 1994c, 1998a, b, 2000, 2001a, b, 2002a, b, 2003a, 2004a, c, d, 2006c, d, 2007a, c, 2008b, c, d, e, f, 2010b, 2011a, b, 2012a, b, 2014c, 2015b, 2017a, b, c, 2018, Niedbała & Buczkowska 2020, Niedbała & Corpuz-Raros 1998, Niedbała & Ermilov 2012, 2013a, b, 2014a, b, 2015, 2017b, 2018a, 2019a, b, 2020, 2022a, b, c, Niedbała et al. 2020, 2021, Niedbała & Osiejuk 2018, Niedbała & Penttinen 2007, Niedbała & Schatz 1996, Niedbała & Starý 2014d, 2015b, e, 2016a, 2022, Niedbała & Szywilewska-Szczykutowicz 2017, Oliveira 2004, Oliveira et al. 2017, Olszanowski et al. 1996, Oudemans 1916a, Pérez-Íñigo 1969a, b, c, Riley 1874, Ryabinin et al. 2018, Schatz 1998, Schmelzle et al. 2008, 2009, 2015, Seniczak et al. 2021, Sergienko 1989, Shimano & Aoki 1997, Subías 2022, Vu 2013, Wang et al. 2002, Weigmann et al. 2015).

Semicosmopolitan, Panpalaearctic

Notes. Subías (2022) recognized Phthiracarus americanus Ewing, 1909 as a subspecies of A. ardua , while Jacot (1938) already synonymized this species with A. ardua .

Subías (2022) treated A. diaphoros (Niedbała,2002) as a subspecies of A. ardua . See note of A. diaphoros . Subías (2022) considered this species as congeneric with Acrotritia munita Niedbała 2006 . However, both species have different shape of trichobothria.

Subías (2022) considered this species as congeneric with Acrotritia paraardua Niedbała et Starý, 2015 . See note of A. paraardua .

Subías (2022) considered this species as congeneric with A. dinota Niedbała et Schatz, 1996 . See note of A. dinota .

Subías (2022) synonymized Acrotritia hauseri (Mahunka, 1991) with A. ardua . See note of A. hauseri .

Subías (2022) synonymized Acrotritia corletti ( Mahunka, 2000) with A. ardua . See note of A. corletti .

ardua affinis Rhysotritia Sergienko, 1989 = Acrotritia ardua C.L. Koch, 1841 )

ardua curticephala Pseudotritia Jacot, 1938 = Acrotritia curticephala ( Jacot, 1938)

ardua jinyunia Rhysotritia Li, Chen et Li, 1990 synonym of Acrotritia sinensis Jacot, 1923

ardua neotropicalis Acrotritia ( Pérez-Íñigo et Baggio, 1993)

Rhysotritia ardua neotropicalis Pérez-Íñigo et Baggio, 1993

Rhysotritia ardua neotropicalis: Niedbała 2004a .

Acrotritia ardua neotropicalis: Subías 2004 , Niedbała & Liu 2018.

Acrotritia neotropicalis: Subías 2022 .

Neotropic: Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Pérez-Íñigo & Baggio 1993, 1997, Subías 2022).

Neotropical, Brazil, endemic ardua penicillata Acrotritia (Pérez-Íñigo, 1969)

Rhysotritia ardua penicillata Pérez-Íñigo, 1969

Acrotritia penicillata ardua: Niedbała & Liu 2018 .

Palaearctic: Spain ( Balogh & Mahunka 1980 a, 1983, Mahunka & Mahunka-Papp 2004, Pérez-Íñigo 1969c, Subías 2022).

Palaearctic, Spain, endemic

Notes. If we accept the suggestion of Mahunka & Mahunka-Papp (2004) that Rhysotritia ardua penicillata Pérez-Íñigo, 1969 is synonym of Rhysotritia ardua hyerogliphica ( Berlese, 1916) , it will be also the synonym of Acrotritia ardua (C.L. Koch, 1841) .

Subías (2022) increased this subspecies to species level, Acrotritia penicillata (Pérez-Íñigo, 1969) .

arilloi Euphthiracarus (Euphthiracarus) Subías, 2009 nov. nom. for Euphthiracarus (Euphthiracarus) reticulatus Niedbała, 2004 , nom. praeoc. for Phthiracarus reticulatus Berlese, 1913 (transferred to genus Euphthiracarus )

Euphthiracarus (Euphthiracarus) arilloi: Subías 2022 .

Neotropic: Mexico, Guatemala ( Niedbała 2004a, Subías 2022).

Neotropical, Mesoamerica armadillo Prototritia ( Berlese, 1916)

Arthrhoplophora (Prototritia) armadillo Berlese, 1916

Arthrhoplophora (Prototritia) armadillo Berlese, 1916 : Niedbała & Liu 2018.

Prototritia armadillo: Hammen 1959 , Mahunka 1980, Mahunka & Mahunka-Papp 1995, Niedbała 2004b. Aedoplophora africana Mahunka, 1977 syn. by Niedbała 2004b.

Prototritia (Prototritia) armadillo: Subías 2022 .

Afrotropic: Kenya, Somalia, Madagascar ( Balogh & Balogh 2002, Berlese 1916, Castagnoli & Pegazzano 1985, Hammen 1959, Mahunka 1977 c, 1980, Mahunka & Mahunka-Papp 1995, Niedbała 2004b, 2008c, 2017a, Niedbała & Starý 2016a, Subías 2022).

Afrotropical, eastern part armatus Notophthiracarus ( Mahunka, 1986)

Hoplophorella armata Mahunka, 1986

Hoplophorella armata: Mahunka 1992 , Niedbała 1992.

Notophthiracarus armatus: Niedbała 2001 a, Niedbała & Liu 2018 .

Phthiracarus (Archiphthiracarus) armatus: Subías 2004 .

Hoplophorella (Hoplophorella) armata: Subías 2022 .

Afrotropic: Tanzania ( Balogh & Balogh 2002, Mahunka 1986, 1992, Niedbała 1992, 2001a, 2017c, Subías 2022).

Afrotropical, Tanzania, endemic asensillata Mesotritia ( Ojeda, 1991)

Perutritia asensillata Ojeda, 1991

Mesotritia (Perutritia) asensillata: Subías 2022 .

Mesotritia asensillata: Niedbała 2004a , Liu & Chen 2010 a, Niedbała & Liu 2018.

Neotropic: Mexico ( Niedbała 2004a, Ojeda 1991, Subías 2022).

Neotropical, Mexico, endemic asiatica Cryptoplophora Gordeeva, Niemi et Petrova-Nikitina, 1998 synonym of Cryptoplophora abscondita Grandjean, 1932

asiatica Oribotritia (Oribotritia) Hammer, 1977

Oribotritia asiatica Hammer, 1977

Oribotritia asiatica: Niedbała 2000, 2011b , 2012b, Subías 2004, Liu et al. 2011a, Liu 2015a, 2021b, Niedbała & Liu 2018.

Oribotritia (Oribotritia) asiatica: Subías 2022 .

Oribotritia asiatica asiatica: Subías 2004 .

Oribotritia (Oribotritia) asiatica asiatica: Subías 2022 .

Oribotritia koreense Mahunka, 1982 syn. by Niedbała 2000.

Palaearctic: Pakistan, Russia (Eastern Siberia), North Korea. Orient: China (Hubei) ( Hammer 1977, Liu 2015a, 2021b, Mahunka 1982c, Niedbała 2000, 2011b, 2012b, 2015b, Subías 2022).

South-Eastern Palaearctic and North-Eastern Oriental, disjunctive, reported from Pakistan, Eastern Siberian, Manchurian areas and from Northern China region asiatica norikoae Oribotritia (Oribotritia) Suzuoka, 1983 subspecies inquirenda

Oribotritia asiatica norikoae: Niedbała 2004c, 2011b , 2012b, Subías 2004, Niedbała & Liu 2018. Oribotritia (Oribotritia) asiatica norikoae: Subías 2022 .

Palaearctic: Japan ( Niedbała 2004c, 2011b, 2012b, Subías 2022, Suzuoka 1983).

Eastern Palaearctic, Japan, endemic

Note. It is difficult to analyze the morphological details in photos 7 and 8 provided by Suzuoka (1983). Therefore, we should agree with Subías (2022) that it is a subspecies inquirenda.

askewi Terratritia Ramsey et Sheals, 1969

Terratritia askewi: Niedbała 2000 , Niedbała & Liu 2018, Subías 2022.

Orient: Malaysia (Sabah) ( Balogh & Balogh 2002, Niedbała 2000, Ramsay & Sheals 1969, Subías 2022). Oriental, Malaysia, endemic aspera Indotritia Niedbała, 2000 synonym of Indotritia (Indotritia) undulata Bayoumi et Mahunka, 1979 aspergillus Steganacarus (Steganacarus) Niedbała, 2004

Steganacarus (Steganacarus) aspergillus: Niedbała & Liu 2018 .

Atropacarus (Rafacarus) aspergillus: Subías 2022 .

Neotropic: Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).