Grammacephalus Haupt

Shah, Bismillah, Naveed, Hassan & Duan, Yani, 2019, Taxonomic review of the leafhopper genus Grammacephalus Haupt (Hemiptera: Cicadellidae: Deltocephalinae: Scaphoideini) with description of a new species from Pakistan, Zootaxa 4688 (1), pp. 71-85: 72-73

publication ID

https://doi.org/10.11646/zootaxa.4688.1.3

publication LSID

lsid:zoobank.org:pub:6A91548F-0C20-4711-950A-11ED5B007BA1

DOI

http://doi.org/10.5281/zenodo.3691801

persistent identifier

http://treatment.plazi.org/id/03B287C3-FF8B-FF8F-FF65-C478FA79F0B5

treatment provided by

Plazi

scientific name

Grammacephalus Haupt
status

 

Key to species of Grammacephalus Haupt 

The following species were described based on female specimens and are therefore omitted from the key: G. minabicus  and G. niveimarginatus  .

1. Male pygofer process absent ( Fig. 3BView FIGURE 3)................................................ Grammacephalus genoicus 

- Male pygofer process present ( Figs 1BView FIGURE 1, 2DView FIGURE 2, 4BView FIGURE 4, 5EView FIGURE 5, 6View FIGURE 6 C–F, 8B, 9C, 10C, 12A)..................................... 2

2. Male pygofer process serrate ( Figs 6View FIGURE 6 C–F, 10C).............................................................. 3

- Male pygofer process not serrate ( Figs 1BView FIGURE 1, 2DView FIGURE 2, 4BView FIGURE 4, 5EView FIGURE 5, 8BView FIGURE 8, 9CView FIGURE 9, 12AView FIGURE 12)............................................ 4

3. Apex of subgenital plate notched, dorsally upturned ( Fig. 6GView FIGURE 6); aedeagal shaft with an apical keel in lateral view ( Fig. 6IView FIGURE 6)..................................................................................... Grammacephalus kempi 

- Apex of subgenital plate entire flat ( Fig. 10DView FIGURE 10); aedeagal shaft with median expansion laterally, without apical keel in lateral view ( Fig. 10FView FIGURE 10).................................................................... Grammacephalus raunoi 

4. Male pygofer process with bifurcated apex ( Figs 2DView FIGURE 2, 12A, BView FIGURE 12).................................................. 5

- Male pygofer process without apical bifurcation ( Figs 1BView FIGURE 1, 4BView FIGURE 4, 5EView FIGURE 5, 8BView FIGURE 8, 9CView FIGURE 9)....................................... 6

5. Subgenital plate with lateral margins slightly incurved at midlength ( Fig. 2FView FIGURE 2); aedeagal shaft strongly reflexed basally with pair of short dorsal processes at apex ( Figs 2H, IView FIGURE 2)........................................... Grammacephalus furcatus 

- Subgenital plate with lateral margins not incurved at midlength ( Fig. 12DView FIGURE 12); aedeagal shaft moderately reflexed basally without appendages ( Figs 12View FIGURE 12 E–G)................................................... Grammacephalus punjabensis  sp.n.

6. Aedeagal shaft with subapical curve in lateral view ( Fig. 4CView FIGURE 4)................................. Grammacephalus pugio 

- Aedeagal shaft without subapical curve in lateral view ( Figs 1CView FIGURE 1, 4EView FIGURE 4, 5LView FIGURE 5, 8GView FIGURE 8, 9FView FIGURE 9)................................... 7

7. Aedeagal shaft not reflexed in lateral view ( Figs 4EView FIGURE 4, 9FView FIGURE 9)...................................................... 8

- Aedeagal shaft somewhat reflexed in lateral view ( Figs 1CView FIGURE 1, 5LView FIGURE 5, 8GView FIGURE 8)............................................. 9

8. Male pygofer process pointed with long subapical spur on dorsal surface ( Fig. 4DView FIGURE 4); aedeagal shaft long and narrowly bladeshaped in lateral view ( Fig. 4EView FIGURE 4)...................................................... Grammacephalus harpago 

- Male pygofer process without subapical spur dorsally ( Fig. 9CView FIGURE 9); aedagal shaft tubular and tapered towards apex ( Fig. 9F, GView FIGURE 9)................................................................................ Grammacephalus rahmani 

9. Aedeagal shaft strongly reflexed basally, broadened and rounded apically ( Figs 8G, HView FIGURE 8).......... Grammacephalus pallidus 

- Aedeagal shaft not strongly reflexed basally (1C, 5L)........................................................ 10

10. Male pygofer process with slight inward curve near apex ( Fig. 5EView FIGURE 5); style apophysis digitate (Fig. H); aedeagal shaft apex acutely tapered in posterior view ( Figs 5KView FIGURE 5).............................................. Grammacephalus indicus 

- Male pygofer process with slight ventral curve near apex ( Fig. 1BView FIGURE 1); style apophysis smooth ( Fig. 1EView FIGURE 1); aedeagal shaft bifid apically in posterior view ( Fig. 1DView FIGURE 1).................................................... Grammacephalus acuticeps