Triplocania einsteini, González-Obando & Carrejo-Gironza & García, 2021

González-Obando, Ranulfo, Carrejo-Gironza, Nancy & García, Alfonso N., 2021, New species of Triplocania Roesler (Psocodea: ‘ Psocoptera’: Ptiloneuridae) from Colombia and Peru, Zootaxa 5080 (1), pp. 1-63 : 26-29

Triplocania einsteini n. sp.

( Figs 61–72 )

Diagnosis. It belongs in species group lucida of Subgroup MPB2A (see Classification below). It is related to T. caquetensis n. sp., from which it differs in the hypandrium: the lateral sclerites are rhomboid, not elongate as in the latter, and the central sclerite has the mesal lateral process larger and stouter than in the former, the median posterior processes have the inner borders slightly concave, and the outer borders mesally convex. The anterior, large endophallic sclerite has the median posterior process broader and shorter than in T. caquetensis , with posterolateral long, slender, distally acuminate processes directed postero-laterally. The lateral endophallic sclerites are also club-shaped, but the clubs are much larger and the stems are much stouter and slightly curved inward than in T. caquetensis . Although similar to that in T. caquetensis , the mesal endophallic sclerite is slightly smaller, with the anterior and posterior processes shorter and stouter. The external parameres are twice as wide than in T. caquetensis , lacking the mesal denticles on the inner borders, and with the anterior processes of the inner borders much shorter and slightly curved, not sinuous ( Fig. 66 ). IX sternum semioval, anterior border slightly convex, with broad Vshaped pigmentation.

Male. Color (in 80% ethanol). Body brown to cream. Head brown, with dark brown spots pattern ( Fig. 63 ). Compound eyes dark brown, ocelli hyaline, with ochre centripetal crescents; antennae and maxillary palps pale brown. Thorax brown, tergal lobes of meso- and metathorax brown with light brown spots. Thoracic pleura brown, with ochre spots. Legs pale brown; coxae with dark spots basally and apically, femora with small dark spots distally; tibiae and tarsi pale brown. Forewings almost hyaline, with a light brown hue, dark brown spots at the ends of veins R, M, CuA 1, CuA 2, CuP and intersection A 1 -CuP; pterostigma with dark brown bands proximally and distally; light brown spots at the end of cells r 5, m 1, and m 2; veins brown ( Fig. 61 ). Hindwings hyaline, veins brown, with brown spots at ends of R and M veins ( Fig. 62 ). Abdomen cream with ochre spots. Epiproct and paraprocts pale brown. Sclerites of hypandrium brown to cream, lateral sclerites darker than the central sclerite. Endophallic sclerites dark brown.

Morphology. As in diagnosis, plus the following: Head ( Fig. 63 ): H/MxW: 1.58; H/D: 2.28; IO/MxW: 0.58. Vertex slightly emarginate, at the level of the upper edge of the compound eyes. Outer cusp of lacinial tips broad, with four denticles. Mx4/Mx2: 1.25. Forewings ( Fig. 61 ): L/W: 2.50. Pterostigma: lp/wp: 5.0. Areola postica triangular: la/ha: 1.84, CuA 1 gently concave distally. Hindwings ( Fig. 62 ): l/p: 2.97. Hypandrium of three sclerites, setose ( Fig. 65 ). Phallosome ( Fig. 66 ), side struts slender, basally convergent, V-shaped. External parameres distally with pores. Paraprocts oval, with abundant medium sized setae and macrosetae on apical third, at least one apically spatulate macroseta, a macroseta near each sensory field, these with 20 trichobothria on basal rosettes ( Fig. 64 ). Epiproct ( Fig. 64 ) wide, broadly rounded posteriorly, setae and macrosetae as illustrated.

Measurements. FW: 3875, HW: 2762, F: 1025, T: 1675, t1: 700, t2: 88, t3: 152, ctt1: 26, f1: 700, f2: 712, f3: 575, Mx4: 250, IO: 345, d: 290, D: 408, IO/d: 1.19, PO: 0.71.

Female. Color. As in the male. Subgenital plate with deep, hyaline median concavity, pigmented area brown, posterior border pale brown. IX sternum hyaline, with anterior triangular area brown; gonapophyses brown to pale brown.

Morphology. Head ( Fig. 69 ): H/MxW: 1.64, H/D: 2.46; compound eyes large, IO/MxW: 0.67. Vertex as in the male. Outer cusp of lacinial tips broad, with four denticles. Mx4/Mx2: 1.50. Forewings ( Fig. 67 ) as in the male, L/W: 2.47, pterostigma: lp/wp: 4.25; areola postica, al/ah: 1.80. Hindwings ( Fig. 68 ): l/w: 2.79. Subgenital plate ( Fig. 71 ) broad, setose, triangular. Gonapophyses ( Fig. 72 ): v1 slender, elongate, acuminate; v2+3: v2 with 11 macrosetae, distal process sinuous, acuminate, with field of microspines. Paraprocts ( Fig. 70 ) elongate, triangular, with setae and macrosetae, one with spatulate apex as illustrated; sensory fields with 24 trichobothria on basal rosettes. Epiproct ( Fig. 70 ) triangular, with apex rounded, mesal field with three macrosetae, two lateral setae and setae along sides and apex, as illustrated.

Measurements. FW: 4075, HW: 2912, F: 1075, T: 1775, t1: 750, t2: 90, t3: 160, ctt1: 27, f1: 662, f2: 675, f3: 585, f4: 500, f5: 325, Mx4: 270, IO: 400, d: 260, D: 400, IO/d: 1.54, PO: 0.65.

Material studied. Holotype male. COLOMBIA. Caquetá. San Vicente del Caguán, Laureles, Reserva Indígena Altamira , 2º27’50.14”N: 74º55’2.06”W, 917 m. 9–10.III.2019. Led light trap in forest canopy. J. Panche. MUSENUV slide code 29939 . Paratype: 1 female, same data as the holotype. MUSENUV slide code 29940 .

Etymology. This species is dedicated to Albert Einstein, theoretical physicist, Nobel Prize in Physics in 1921, author of the Theory of Relativity.

Figures

FIGURES 61–66. Triplocania einsteini n. sp. Male. 61. Forewing. 62. Hindwing. 63. Front view of head. 64. Epiproct and right paraproct. 65. Hypandrium. 66. Phallosome. Scales in mm.

FIGURES 67–72. Triplocania einsteini n. sp. Female. 67. Forewing. 68. Hindwing. 69. Front view of head. 70. Epiproct and right paraproct. 71. Subgenital plate. 72. Ninth sternum and right gonapophyses. Scales in mm.