Procoilodes adrastus Ocampo, 2002

Ocampo, Federico C., 2002, Procoilodes adrastus Ocampo, A New Genus And Species Of Miocene Hybosorid In Amber From The Dominican Republic (Coleoptera: Scarabaeoidea: Hybosoridae), The Coleopterists Bulletin 56 (1), pp. 123-126 : 125-126

Procoilodes adrastus Ocampo , new species

( Figs. 1 –4)

Type Material. Holotype from the Dominican Republic (specimen number AMNH DR­10–136 ) embedded in Miocene amber . Holotype deposited in the Amber Fossil Collection , Dept. of Entomology , American Museum of Natural History, New York .

Description. Holotype. Length 4.73 mm; greatest width 2.76 mm. Color dark brown. Head. Frons with surface heavily and finely tuberculate, vertex with 2 conical tubercles. Eye canthus elongate, rectangular, apex truncate. Frontoclypeal suture not visible. Clypeus with surface similar to that of frons; anterior margin slightly rounded, lateral margins arcuate and narrowly reflexed. Eyes subglobose. Mandibles large; external edge broadly rounded; dorsal surface rugose, concave; subapex with well­developed, dorsal tooth. Maxilla and galea with dense setae; maxillary palpus with 4 segments. Labium with sparse, long setae. Pronotum. Surface weakly convex, heavily and finely tuberculate, slightly depressed on middle in anterior half. Edges of anterior angles, lateral margin, and sides of posterior margin smooth. Scutellum . Shape subtriangular; apex acute. Elytra. Surface shagreened, with 1 stria next to elytral suture. Humeral umbone elevated. Legs. Protibiae (Fig. 3) on external margin with 3 long, acute teeth and with 2 denticles between both apical and median and between median and basal teeth, and with 5 denticles behind basal tooth; apical spur not reaching apex of second tarsal segment. Protarsus with segments 1–4 subequal in length and each shorter than fifth, each with slender distal setae. Meso­ and metatarsi with segments 1 and 5 longer than any of segments 2– 4, segments 1–4 with slender distal setae.

Remarks. The tree that produced the resin that trapped this specimen is Hymenaea protera Poinar (Leguminosae, Caesalpinioidea). The amber in which it is embedded was dated as Miocene in age (Grimaldi 1995).

Etymology. From the classical mythological name ‘‘ Adrastus,’’ which means ‘‘he who holds his position’’ or ‘‘the unescapable,’’ a name perfectly applied to this specimen which has been embedded and immovable in amber for more than 15 million years.

126 THE COLEOPTERISTS BULLETIN 56(1), 2002

I thank David Grimaldi ( American Museum of Natural History , NY) for

making the specimen available to me for study. I am grateful to Brett Ratcliffe and Mary Liz Jameson (University of Nebraska, Lincoln, NE) for their guidance in preparing the manuscript. Two anonymous reviewers are thanked for their helpful remarks to improve the manuscript. This project was supported by an NSF/ Biotic Survey and Inventory Grant (DEB 9870202) to B. C. Ratcliffe and an NSF/ PEET grant (DEB 9712447) to B. C. Ratcliffe and M. L. Jameson.

Figures

Fig. 1. Holotype of Procoilodes adrastus embedded in amber, dorsal view.