Helicopsyche (Feropsyche) circulata, Johanson, Kjell Arne & Holzenthal, Ralph W., 2004

Johanson, Kjell Arne & Holzenthal, Ralph W., 2004, Thirteen new species and new distribution records of Helicopsyche (Feropsyche) Johanson from Venezuela (Trichoptera: Helicopsychidae), Zootaxa 711, pp. 1-40 : 25-27

publication ID	https://doi.org/ 10.5281/zenodo.169460
publication LSID	lsid:zoobank.org:pub:5403C332-6597-48D3-BD4D-7AD6D4CDC2CA
DOI	https://doi.org/10.5281/zenodo.6271574
persistent identifier	https://treatment.plazi.org/id/03B1879D-FFAF-DE50-FE98-FB0C7DB7FB43
treatment provided by	Plazi
scientific name	Helicopsyche (Feropsyche) circulata
status	sp. nov.

Treatment

Helicopsyche (Feropsyche) circulata , new species

Fig. 72–78 View FIGURES 72 – 78 , 90 View FIGURE 90

The three species, H. circulata , new species, H. perija , new species, and H. planorboides Machado , all have a characteristically wide and rounded distal part of the gonocoxite. In H. circulata , the anterodorsal margin of the gonocoxite is nearly straight, while in H. planorboides it is strongly concave. Also, tergum X is simple in H. circulata , while in H. planorboides a pair of lateral lobes originate midway on the segment. Helicopsyche circulata can be separated from H. perija by the shorter anterior lobe of segment IX, and the longer sternum VI process.

Male. Head: Antennal scape as long as eye diameter. Maxillary palp segments equally long, each segment as long as eye diameter. Cephalic warts oval, strongly convex, about half as long as eye diameter; with pale grey and golden brown setae. Forewing golden brown, length 4.1–4.3 mm. Sternum VI process ( Fig. 72, 73 View FIGURES 72 – 78 ) nearly 2/3 its segment length, covered by microtrichiae; in lateral view ( Fig. 72 View FIGURES 72 – 78 ), nearly straight, slightly tapering along its length; in ventral view ( Fig. 73 View FIGURES 72 – 78 ), slightly tapering; apex bearing numerous ventral thorny lamellae ( Fig. 73 View FIGURES 72 – 78 ).

Male genitalia ( Fig. 74–78 View FIGURES 72 – 78 ). Segment IX, in lateral view ( Fig. 74 View FIGURES 72 – 78 ), with anterior lobe triangular, oriented anteriad and located midlaterally on segment; anterodorsal margin nearly straight, anteroventral margin concave; in dorsal view ( Fig. 75 View FIGURES 72 – 78 ), with inner margin narrowly ellipsoid; in ventral view ( Fig. 76 View FIGURES 72 – 78 ), with pointed posterior process; lateral apodeme present as anteriorly oriented, slightly curving line ( Fig. 74 View FIGURES 72 – 78 ) reaching anterior margin; submarginal line absent; tergal transverse apodeme absent; sternal transverse apodeme short. Segment X, in lateral view ( Fig. 74 View FIGURES 72 – 78 ), oriented posteroventrad, nearly straight; narrowing along its length toward rounded apex; in dorsal view ( Fig. 75 View FIGURES 72 – 78 ), narrow, with subparallel lateral margins; apex with Vshaped notch ( Fig. 75 View FIGURES 72 – 78 ); with about 11 pairs of about equally long megasetae in longitudinal group starting at proximal half on segment. Superior appendage ( Fig. 74 View FIGURES 72 – 78 ) clubshaped in lateral view, parallelsided in dorsal view ( Fig. 74, 75 View FIGURES 72 – 78 ); oriented laterally ( Fig. 75 View FIGURES 72 – 78 ). Primary branch of gonocoxite, in lateral view ( Fig. 74 View FIGURES 72 – 78 ), widely clubshaped, with slightly undulate dorsal margin; apex rounded in lateral and dorsal views ( Fig. 74, 75 View FIGURES 72 – 78 ); central part of primary branch about as wide as height of central part of tergum X ( Fig. 74 View FIGURES 72 – 78 ); anterodorsal margin nearly straight, smooth; posteroventral margin slightly convex, undulated; basimesal lobe short, rounded in lateral view ( Fig. 74 View FIGURES 72 – 78 ); in ventral view, sharply triangular with setal bases produced posteriorly ( Fig. 76 View FIGURES 72 – 78 ); with nearly straight and parallelsided, smooth median margins; with about 4 long megasetae on apical margin; basal plate not visible in lateral view ( Fig. 74 View FIGURES 72 – 78 ); in ventral view ( Fig. 76 View FIGURES 72 – 78 ), strongly pointing anteriorly. Phallus, in lateral view ( Fig. 77 View FIGURES 72 – 78 ), generally slender, smoothly bent ventrad along its length, dorsal margin sigmoid along proximal 2/3; ventral margin gently concave; anterior 1/5 nearly 2x broader than its central part ( Fig. 77 View FIGURES 72 – 78 ); in ventral view ( Fig. 78 View FIGURES 72 – 78 ), slightly narrowing toward midlength; phallobase present; endotheca weakly produced ( Fig. 77 View FIGURES 72 – 78 ); sperm channel divided into thick posterior and slen der anterior parts; sclerotized posteroventral part narrow ( Fig. 77 View FIGURES 72 – 78 ).

Holotype male: VENEZUELA: Aragua: 1 km E Estación Biológica Rancho Grande, 10.352°N, 67.680°W, 1100 m, 27.i.1994, Holzenthal, Cressa, Gutic ( UMSP 000022205) ( UMSP, pinned).

Paratype: same data as holotype — 1 male ( UMSP, pinned).

Distribution. Venezuela (Aragua).

Etymology. The specific epithet, circulata , is derived from the Greek kirkos, ring, and the suffix atus, provided with, referring to the circular shape of the gonocoxite. The name is to be treated as a noun in apposition.