Goloboffia megadeth, Ferretti & Ríos-Tamayo & Goloboff, 2019

Ferretti, Nelson E., Ríos-Tamayo, Duniesky & Goloboff, Pablo A., 2019, Four new species of Goloboffia (Mygalomorphae: Migidae) from Chile, Zootaxa 4712 (2), pp. 251-268 : 258-260

Goloboffia megadeth sp. nov.

( Figs 23–47 , 71 )

Type material. Holotype: ♂, MACN-Ar 36117, Chile, Región de Coquimbo (IV), Ovalle, Monte Redondo, 15 km north of Mantos de Hornillos , 1.xi.1988, P. Goloboff, E. Maury and C. Szumik leg. Paratype: 1 ♀, MACN-Ar 36131, same collection data .

Additional material examined. 2 ♀♀ and 3 juveniles, MACN-Ar 36131, Chile, Región de Coquimbo (IV), Ovalle, Monte Redondo, 15 km north of Mantos de Hornillos , 1.xi.1988, P. Goloboff, E. Maury and C. Szumik leg.

Etymology. The specific epithet is a noun taken in apposition from the band’s name Megadeth, a U.S. heavy metal band lead by guitarist Dave Mustaine.

Diagnosis. Males of Goloboffia megadeth sp. nov differ from G. vellardi in having a thinner and longer em- bolus with a wavy tip ( Figs 33–36 ); females differ from the other known species by the shape of the spermathecae with large and rounded lobes and very short ducts of the same length as spermathecal head width ( Fig. 47 ).

Description. Holotype male, MACN-Ar 36117. Coloration (in alcohol): Carapace and chelicerae reddish brown, legs brown, abdomen brownish ( Figs 23–27 ). Total length 8.18; carapace 2.55 long, 2.53 wide. Fovea deep, straight with recurved ends, 0.51 wide. Ocular group 1.28 wide, occupying 0.51 of cephalic width ( Fig. 25 ). Eye sizes and interdistances: AME 0.13, PME 0.12, ALE 0.33, PLE 0.13, AME–AME 0.12, AME–ALE 0.24, PLE–PME 0.10, PME–PME 0.52. Labium 0.49 long, 0.55 wide, without cuspules. Labium separated from sternum by shallow groove. Maxillae without cuspules ( Fig. 28 ). Fangs keeled and chelicerae with subequal teeth, 3 promarginal and 3 retromarginal, and 9 denticles between margins. Sternum elongated 1.79 long, 1.47 wide. Posterior sigillae deep, airfoil shaped ( Fig. 24 ). Leg formula 1423. Segment lengths (femur, patella, tibia, metatarsus, tarsus, total): I: 2.97, 1.32, 2.16, 1.92, 0.78, 9.15. II: 2.28, 1.03, 1.93, 1.64, 0.43, 7.31. III: 2.17, 0.98, 1.47, 1.38, 0.71, 6.71. IV: 2.46, 1.26, 2.37, 1.25, 0.73, 8.07. Palp: 1.6, 0.37, 1.76, –, 0.5, 4.23. Spines: Femur: Palp: 1d; I: 1 – 1 – 1d, 1 – 1 – 1 – 1 – 1 – 3p, 1r; II: 1 – 1 – 1d, 1 – 1 – 1 – 1p; III: 1 – 1d, 1 – 1 – 1p; IV: 1 – 1d. Patellae: Palp: 0; I: 1p, 5v; II: 1p, 2v; III: 2 – 2 – 2 – 3d, 3r; IV: 23d. Tibia: Palp: 0; I: 1 – 1 – 1p, 2 – 1 – 2 – 2v, 1 – 1 – 1 – 1r ( Fig. 29 ); II: 2 – 1 – 2 – 2v, 1 – 1 – 1p; III: 0; IV: 0. Metatarsus: I: 1 – 1 – 1 – 1p, 1 – 1 – 1r ( Fig. 30 ); II: 1 – 1 – 1 – 1p, 1 – 1 – 1 – 1r; III: 0 ( Fig. 31 ); IV: 0 ( Fig. 32 ). Tarsus: I – IV: 0. Bulb globose on base with long and thin embolus wavy at tip ( Figs. 33–34 ).

Female (MACN-Ar 36131), coloration (in alcohol): Carapace, legs and chelicerae brown, abdomen light gray ( Figs. 37–41 ). Total length 9.35; carapace 3.77 long, 3.22 wide. Fovea deep, straight with recurved ends, 0.51 wide. Ocular group 1.62 wide, occupying 0.61 of cephalic width ( Fig. 42 ). Eye sizes and interdistances: AME 0.12, PME 0.15, ALE 0.40, PLE 0.14, AME–AME 0.16, AME–ALE 0.36, PLE–PME 0.09, PME–PME 0.63. Labium 0.62 long, 0.52 wide, with 8 cuspules. Labium separated from sternum by shallow groove. Maxillae with 7/9 (right/left) cuspules ( Fig. 43 ). Fangs keeled and chelicerae with subequal teeth, 4 promarginal and 4 retromarginal, and 11 denticles between margins. Sternum elongated, 1.79 long, 1.65 wide. Posterior sigillae deep, large and airfoil shaped, shallow ( Fig. 38 ). Leg formula 4123, tibiae and metatarsi I and II flattened, with very long spines ( Figs. 44–45 ). Segment lengths (femur, patella, tibia, metatarsus, tarsus, total): I: 2.41, 1.43, 1.69, 1.18, 0.53, 7.24. II: 2.2, 1.02, 1.33, 1.05, 0.5, 6.1. III: 1.7, 1.01, 1.17, 0.86, 0.52, 5.26. IV: 2.47, 1.42, 1.79, 2.01, 0.45, 8.14. Palp: 1.39, 0.79, 0.94, –, 0.70, 3.82. Spines: Femur: Palp and legs I – IV, 0. Patella: Palp: 1p; I: 1p; II: 0; III: 25p; IV: 123d. Tibia: Palp: 1p; I: 4p, 10r; II: 4p, 6r; III: 6p; IV: 0. Metatarsus: I: 4p, 6r, 4v; II: 5p, 4r; III: 3p, 6d, ventral comb of 10 large and 14 small spines ( Fig. 46 ); IV: ventral comb of 17 large spines. Tarsus: Palp: 3p, 3r, 6v; I: 3p, 4r, 14v; II: 3p, 5r, 13v; III: 12v; IV: 0.

Spermathecae with rounded lobes pointed outwards and very short ducts (same length as spermathecal head width), constriction between ducts and lobes well developed ( Fig. 47 ).

Distribution. Known only from the type locality ( Fig. 71 ).

Figures

FIGURE 71. Known distribution of Goloboffia species in Chile.

FIGURES 23–28. Goloboffia megadeth sp. nov., holotype male, MACN-Ar 36117. 23. Carapace, dorsal view. 24. Labium and sternum. 25. Eye group. 26. Abdomen, dorsal view. 27. Abdomen, ventral view. 28. Labium and maxillae. Scale bars=1 mm.

FIGURES 29–32. Goloboffia megadeth sp. nov., holotype male, MACN-Ar 36117. 29. Tibia I, ventral view. 30. Metatarsus and tarsus I, ventral view. 31. Metatarsus and tarsus III, ventral view. 32. Metatarsus and tarsus IV, ventral view. Scale bars=1 mm

FIGURES 33–36. Goloboffia megadeth sp. nov., holotype male, MACN-Ar 36117. 33. Palpal bulb, dorsal view. 34. Palpal bulb, ventral view. 35. Palp, retrolateral view. 36. Palp, prolateral view. Scale bars=1 mm.

FIGURES 37–42. Goloboffia megadeth sp. nov., paratype female, MACN-Ar 36131. 37. Carapace, dorsal view. 38. Labium and sternum. 39. Carapace, lateral view. 40. Abdomen, dorsal view. 41. Abdomen, ventral view. 42. Eye group. Scale bars=1 mm.

FIGURES 43–47. Goloboffia megadeth sp. nov., paratype female, MACN-Ar 36131. 43. Labium and maxillae. 44. Metatarsus I, prolateral view. 45. Tibia I, prolateral view. 46. Tarsus III, retrolateral view. 47. Spermathecae, dorsal view. Scale bars=1 mm.