Rhamphomyia (Holoclera) portugalica, Barták & Kubík, 2012

Barták, Miroslav & Kubík, Stepán, 2012, A review of the Palaearctic species of Rhamphomyia subgenus Holoclera (Diptera: Empididae) with description of 5 new species, Revue suisse de Zoologie 119 (3), pp. 385-407 : 395-397

Rhamphomyia (Holoclera) portugalica sp. n. Fig. 4

HOLOTYPE: CULSP; 1 Ƌ; Portugal, 5 km N of Formalicao , 40°28´31´´N, 7°21´32´´W, 930 m; Castanea wood, sweeping vegetation; 23.v.2008; M. Barták.

PARATYPES: CULSP; 21 ♀; same data as the Holotype .- 1 ♀; Serra da Estrella , forest, 40°24´13´´N, 7°35´10´´W, 1450 m; 16.-17.vii.2009; M. Barták .

DISTRIBUTION: Portugal.

DATE OF OCCURRENCE: May-July.

DESCRIPTION

Male: Eyes holoptic, facets in ventral half of eye smaller than in dorsal half. Frons (small area just above antenna and below ocellar triangle) brown, brown microtrichose, setae absent. Ocellar triangle with 2 pairs of short setae, ocellars broken in the single male at hand. Face brown, microtrichose, 0.10 mm broad ventrally and about 0.09 mm long, without setae. Occiput brown, microtrichose, sparsely black setose (setae arranged in two almost complete and regular rows). Antenna brownish black, length of antennal segments (in 0.01 mm scale) = 9: 7: 22: 11, both basal antennomeres short setose (the longest setae about 0.06 mm long). Labrum brown, 1.5 times as long as head is high. Palpus brown, only 1 short seta on base of broadened terminal part and further three setae on base. Gena extremely narrow, clypeus microtrichose. Thorax brown, microtrichose, mesoscutum light brownish grey viewed from the front or from Rhamphomyia (Holoclera) portugalica sp. n., male terminalia (macerated), lateral view. Scale 0.1 mm.

sides and uniformly dark almost velvety brownish black in dorsal view and with two narrow somewhat lighter stripes between rows of dorsocentrals in caudal view. All thoracic setae black. Chaetotaxy: 2 setae on proepisternum; both prosternum and proepisternal depression without setae; 6-8 uniserial dorsocentrals (about 0.15 mm long in middle of their rows), last 1-2 (prescutellar) pairs slightly longer (but one shorter seta present on both sides close to scutellum); acrostichals absent; intrahumeral absent; one strong posthumeral (no additional setae laterad of dorsocentrals in presutural part of mesoscutum); 1 long postpronotal and 2 very short additional setae; 2 notopleurals (1-2 shorter setae on anterior part of notopleuron); 1 rather long supraalar (inserted in cranial position, almost in prealar region); 1 strong and 2 very small postalars; 4 scutellars; laterotergite with black setae. Legs including coxae brown, microtrichose (hind femur lustrous except for the tip), black setose. One very short and poorly distinct seta in posteroapical comb on hind tibia. Fore femur with short setation (no prominent setae), anterior row of spine like (sensory) setae about 0.02 mm long. Fore tibia with short setation only, without prominent setae (except preapical ventral). Mid femur with short setation, no prominent setae. Mid tibia with one anteroventral seta in apical third subequally long as remaining short setation. Hind femur with short setation, without prominent setae. Hind tibia slightly broadened apically, with 1-3 posterodorsal setae slightly stronger but no longer than remaining short setation. Basal tarsomeres of both fore and mid legs thin and short setose, basal tarsomere of hind leg thin, short setose, ventrally with several slightly prominent setae. Measurements (in 0.01 mm scale): length: f1 (75), f2 (95), f3 (128), t1 (75), t2 (80), t3 (125), bt1 (50), bt2 (40), bt3 (60), width: bt1 and bt2 (3), bt3 (6). Wing light brownish, stigma brown, veins brown, anal vein (A1) depigmented in middle part, but apical part distinct. Costal seta long, axillary angle slightly obtuse. M2/D = 2.2, CuA1 ratio = 5.0, lw: ww = 2.6. Halter yellowish brown with darker stem, calypter light brown with dark fringes. Abdomen brown, brown microtrichose, black setose. Hind marginal setae on sides of tergites 2-3 subequally long as their segments, on remaining segments much shorter, discal setae slightly shorter than marginals, dorsum of tergites with short setae, 1st sternite with 2 setae. The 8th tergite and sternite scarcely separated. Terminalia ( Fig. 4 ): hypandrium narrow, with a single long submedian seta, and scarcely differentiated from sclerotized membrane; epandrium approximately rectangular with elongate dorsoapical corner; cercus unusually shaped: trilobate, two dorsal lobes visible in lateral view, lower of them broader and much longer than upper one and cercus with the third short internal lobe visible after dissection; phallus extremely thin to the tip. Length of body 2.9 mm, wing 3.0 mm.

Female: Similar to male but with the following differences. Eyes dichoptic, facets subequal in size. Frons brown, microtrichose, 0.18-0.23 mm long and 0.09 mm broad ventrally and 0.08 mm broad dorsally (slightly narrowing above), with 4-5 short setae on each side (about 0.05 mm). Ocellar setae half as long as the distance between front ocellus and base of antenna. Labrum 1.5-1.8 times as long as head is high. Fore coxa paler than in male, yellowish brown anteriorly about base. Legs as in male, setae on mid and hind tibia more distinct, mid tibia often with 1-2 anterodorsal setae, hind femur often with 1-3 ventral setae on apical third. Basal tarsomeres with more distinct ventral setae. Measurements (in 0.01 mm scale): length: f1 (67-85), f2 (85-105), f3 (123-150), t1 (70-90), t2 (80-100), t3 (94-140), bt1 (37-50), bt2 (32-40), bt3 (42-63), width: bt1 and bt2 (6), bt3 (9). M2/D = 1.6-2.1, CuA1 ratio = 3.7-5.3, lw: ww = 2.6- 3.1. Abdomen brown, microtrichose. Hind marginal setae on sides of tergites 2-4 half as long as their corresponding segments, on remaining segments much shorter. Length of body 2.7-4.2 mm, wing 2.6-3.7 mm.

DIFFERENTIAL DIAGNOSIS: R. (Holoclera) portugalica sp. n. belongs to the R. (Holoclera) nigripennis group. However, it differs from other species of this group by its long labrum and unusually (trilobate) shaped male cercus. Females should be compared with R. (Holoclera) tenuipes Becker and R. (Holoclera) lamellata Collin (both having fore coxae paler than pleurae and supraalar seta present), but may be differentiated according to the key.

DERIVATIO NOMINIS: the specific name is derived from the country of origin

( Portugal).

Figures

FIG. 4