Turanophlebia martynovi Pritykina, 1968

Fleck, Günther, Bechly, Günter, Martínez-Delclòs, Xavier, Jarzembowski, Edmund A. & Nel, André, 2004, A revision of the Upper Jurassic-Lower Cretaceous dragonfly family Tarsophlebiidae, with a discussion on the phylogenetic positions of the Tarsophlebiidae and Sieblosiidae (Insecta, Odonatoptera, Panodonata), Geodiversitas 26 (1), pp. 33-60 : 41-42

Turanophlebia martynovi Pritykina, 1968 ( Fig. 3 )

Turanophlebia martynovi Pritykina, 1968: 43 , 44, textfig. 14, pl. 3, fig. 4. — Carpenter 1992: 73, fig. 46.4. — Nel et al. 1993: 89, 90, fig. 70.

HOLOTYPE. — Specimen No. 2554/21 ( PIN).

GEOLOGICAL SETTING. — Upper Jurassic, Callovian- Kimmeridgian or Oxfordian-Kimmeridgian ( Zherikhin & Gratshev 1993; Mostovski & Martínez- Delclòs 2000), Karatau, Chimkent region, Southern Kazakhstan, C.I.S.

DIAGNOSIS. — T. martynovi differs from T. sibirica and T. anglicana n. sp. in the following character states: pterostigma covering six cells; arculus opposite Ax2; two rows of cells between C and RA distal of pterostigma. T. martynovi has two rows of cells in hind wing anal area, unlike T. neckini n. comb. T. martynovi has a strong angle in its hind wing arculus; its subdiscoidal space is divided into three small cells, unlike other Turanophlebia species. It differs from T. vitimensis n. sp. in the presence of only six rows of cells in cubito-anal area between CuA and posterior wing margin. It differs from T. mongolica n. sp. in the presence of its vein “O” oblique.

? Turanophlebia sibirica Pritykina, 1977 Turanophlebia sibirica Pritykina, 1977: 84 , 85, textfig. 2, pl. 1, fig. 2. — Nel et al. 1993: 91.

HOLOTYPE. — Specimen No. 1289/1258 ( PIN).

GEOLOGICAL SETTING. — Lower Cretaceous, Zaza Formation, Neocomian or Barremian-Aptian ( Zherikhin et al. 1999; Mostovski & Martínez-Delclòs 2000). Baissa, Vitim river, Transbaikalia, C.I.S.

DIAGNOSIS. —? T. sibirica differs from T. martynovi in the presence of only one row of cells in the area between C and RA distal of pterostigma. It differs from T. anglicana n. sp. in its longer wings. It differs from T. neckini n. comb. in the presence of two or three rows of cells in the hind wing anal area.

REMARKS

The type specimen of? T. sibirica is very poorly preserved and incomplete. It is nearly impossible to be accurate of its attribution to the genus Turanophlebia : its cubito-anal area is too fragmentary to determine the exact number of cells rows between CuA and posterior wing margin (Pritykina’s wing reconstruction is optimistic); its vein IR1 is badly preserved. The only argument is the large number of cells in the apical part of its wing.

Turanophlebia neckini ( Martynov,1927) n. comb. ( Fig. 4 )

Tarsophlebia neckini Martynov, 1927: 757 , 758, figs 1, 2. — Nel et al. 1993: 87.

HOLOTYPE. — Specimen No. 2452/3 ( PIN).

GEOLOGICAL SETTING. — Upper Jurassic, Callovian- Kimmeridgian or Oxfordian-Kimmeridgian ( Zherikhin & Gratshev 1993; Mostovski & Martínez- Delclòs 2000), Karatau, Chimkent region, Southern Kazakhstan, C.I.S.

DIAGNOSIS. — T. neckini n. comb. differs from the other Turanophlebia species in the presence of only one or two rows of cells in the hind wing anal area.

REDESCRIPTION

There is some imprecision in the original description of Martynov (1927), thus a redescription is necessary. Imprint of a hind wing, with the distal half partly destroyed. Wing probably hyaline, pterostigma missing; wing about 39.0 mm long; 7.3 mm wide; distance from base to arculus, 6.0 mm; from arculus to nodus, 10.2 mm; median and submedian areas free of cross-veins; CuP strongly curved, nearly opposite Ax2, basally closing subdiscoidal space; primary antenodal braces Ax1 and Ax2 stronger than secondary antenodal cross-veins, without cross-veins between them, 2.0 mm apart; Ax1 is 3.8 mm from wing base; arculus opposite Ax2; 14 secondary antenodal cross-veins distal of Ax2, not aligned with the cross-veins of second rank between ScP and RA; numerous cross-veins in the area between RA and RP, between arculus and nodus; a long “gap” without cross-veins between arculus and RP3/ 4 in the area between RP and MA; MP + CuA strongly curved just before its fusion with MAb; a sharp angle between MP + CuA and MAb; presence of a long fusion between MAb and MP + CuA before CuA separates from MP, 1.1 mm long, characteristic of the Tarsophlebiidae (Nel et al. 1993) ; RP + MA, MA and MAb, MP + CuA + MAb, and basal free part of CuA well aligned in arculus, as in other Tarsophlebiidae (Nel et al. 1993) ; discoidal space basally opened; presence of a two-celled “tarsophlebiid pseudo-discoidal space” just distal of MAb in postdiscoidal area; subdiscoidal area divided into two cells by a cross-vein; AA without any strong posterior branches; anal area with one or two rows of cells; posterior wing margin round- ed; a short petiole, 1.4 mm long; AA reaching free part of CuA at sharp angle; no CuAb (sensu Fleck et al. 2003); CuA without strong posterior branches; six rows of small cells in cubito-anal area; CuA reaching posterior wing margin just distal of nodus level; area between MP and CuA with one row of cells in its basal part but greatly widened in its distal half; postdiscoidal area with two rows of cells in its basal part, narrowed in its mid part and slightly widened distally; bases of RP3/4 and IR2 between arculus and nodus, midway between arculus and nodus, 5.2 mm from nodus; apparent base of IR2 on RP3/4; nodal Cr and subnodus strongly oblique; base of RP2 aligned with subnodus; oblique vein “O” three small cells distal of base of RP2; numerous Bq crossveins, but apparently no cross-vein in basal part of area between RA, RP, RP3/4 and IR2; numerous postnodal cross-veins between C and RA, not aligned with the postsubnodal cross-veins; base of IR1 about 10 cells distal of that of RP2; IR1 well defined, long, not zigzagged and only slightly curved; one row of cells between RP1 and IR1; area between RP3/4 and MA distally widened.

DISCUSSION

This fossil has the diagnostic characters of the genus Turanophlebia , i.e. cubito-anal area broad, with six rows of cells between CuA and posterior wing margin, more than 25 postnodal crossveins, more than 10 secondary antenodal crossveins in hind wing, IR1 longer than in Tarsophlebia eximia . Thus, we propose to transfer it into the genus Turanophlebia .

Figures

FIG. 3. — Turanophlebia martynovi Pritykina, 1968, holotype (PIN 2554/21). Scale bar: 10 mm.

FIG. 4. — Turanophlebia neckini (Martynov, 1927) n. comb., holotype (PIN 2452/3). Scale bar: 10 mm.