Sphegina, Meigen, 1822

Steenis, Jeroen Van, Hippa, Heikki & Mutin, Valeri A., 2018, Revision of the Oriental species of the genus Sphegina Meigen, 1822 (Diptera: Syrphidae), European Journal of Taxonomy 489, pp. 1-198 : 7-17

publication ID

https://doi.org/ 10.5852/ejt.2018.489

publication LSID

lsid:zoobank.org:pub:B234C53F-BB56-4F3D-B400-F214E19DCF90

persistent identifier

https://treatment.plazi.org/id/03AF879D-1926-FFF7-6A8B-A256755BFA52

treatment provided by

Valdenar

scientific name

Sphegina
status

 

Key to the Oriental species of Sphegina View in CoL

1. Sternum I quadratic to oval in shape, pilose; postmetacoxal bridge narrow, arcuate or with large dorso-medial subtriangular incision. (Subgenus Sphegina Meigen, 1822 View in CoL ) ..................................... 2

– Sternum I narrow lanceolate, several times as long as wide or membranous, non-pilose; postmetacoxal bridge wide, its dorsal margin almost straight, at most with small subtriangular medial incision. (Subgenus Asiosphegina Stackelberg, 1974 View in CoL ) ................................................................................... 7

2. Scutellum with 4 long setae at posterior margin; occiput on postero-lateral side of ocellar triangle with very long and strong black pile ( Fig. 3E View Fig ) .......................... S. (S.) quadriseta Huo & Ren, 2006

– Scutellum with 2 long setae at posterior margin; pile on occiput rather short, of even length throughout ( Fig. 3C View Fig ) ...................................................................................................................................... 3

3. Tergum III black, with yellow anterior fascia ........................................... S. (S.) guptai Mutin, 1998 View in CoL – Tergum III entirely dark brown to black ........................................................................................... 4

4. Terga of nearly uniform width, only slightly widening towards posterior end of tergum IV; frontal prominence rather strongly protruding ( Fig. 3C View Fig ) ............................................................................. 5

– Terga widening from posterior end of tergum II towards posterior end of tergum III, then narrowing towards posterior end of tergum IV; frontal prominence very strongly protruding ......................... 6

5. Ventral part of face yellow, dorsal part greyish-black; metatibia of nearly uniform width throughout ( Fig. 19A View Fig ); genitalia, Fig. 1 View Fig B–D ................................................................ S. (S.) abbreviata sp. nov.

– Face entirely yellow ( Fig. 3C View Fig ); metatibia clearly widened at apical 1/5, club shaped ( Fig. 19B View Fig ); genitalia, Fig. 4 View Fig B–D .................................................................................... S. (S.) angustata sp. nov.

6. Dorsal line of frontal prominence in profile straight, ending just before the ocellar triangle; mouth edge strongly projecting, vibrissal angle more strongly protruding than the frontal prominence ........................................................ S. (S.) uncinata Hippa, van Steenis & Mutin, 2015

– Dorsal line of frontal prominence in profile convex, ending clearly before the ocellar triangle; mouth edge less strongly projecting, vibrissal angle equally or less protruding than the frontal prominence ................................................................................... S. (S.) kumaoniensis Mutin, 1998

7. Tergum I with a horizontal row of 2–4 strong black to yellow setae at lateral margin (e.g., Figs 13 View Fig B– C, E, 47B–C, F), in some specimens only one seta is present .......................................................... 8

– Tergum I with an oblique, sometimes almost transverse row of 3–5 dark brown to light yellow setae or seta-like pile deviating from the other vestiture on the posterolateral part (e.g., Figs 13A, D View Fig , 47A View Fig , D–E) or without marginal seta-like pile ......................................................................................... 62

8. Sternum IV posteriorly with conspicuous long or short, often dark brown to black, setae in an asymmetrical pattern, deviating strongly from other vestiture (e.g., Figs 16A View Fig , 17A View Fig , 27A View Fig , 37B View Fig , 42B View Fig ); surstyli strongly asymmetric (e.g., Figs 16A View Fig , 27C, E View Fig , 37C, E View Fig ), in exceptional cases almost symmetrical (e.g., Fig. 42C, E View Fig ) ......................................................................................................... 9

– Sternum IV posteriorly at most with groups of longer light yellow seta-like pile in a symmetrical pattern, deviating only slightly from other vestiture, (e.g., Figs 7A View Fig , 8A View Fig , 12A View Fig , 14A View Fig , 18A View Fig ); surstyli almost symmetrical (e.g., Figs 7B, D View Fig , 8B, D View Fig , 12B, D View Fig , 14B, D View Fig , 18 View Fig B–C) ........................................ 34

9. Wing with infuscate pattern at transverse veins and apex (e.g., Figs 10E View Fig , 15D, F View Fig ) ....................... 10 – Wing hyaline, without infuscate pattern (e.g., Figs 15A View Fig , 28C View Fig , 35C View Fig ) .............................................. 17

10. Sternum I with 2–3 black setae at lateral margin, placed posteriorly ( Fig. 13B View Fig ); metafemur with 1–3 long sub-apical setae antero-dorsally (sometimes these setae are broken off and the sockets may be very hard to see) ( Figs 52A View Fig , 72 View Fig E–F); wing vein R 2+3 subapically without an appendix (e.g., Figs 10E View Fig , 41E View Fig , 50B View Fig ) .........................................................................................................................................11

– Sternum I with 2 black setae at lateral margin which are very widely separated, one placed anteriorly the other posteriorly ( Fig. 13C View Fig ); metafemur without long sub-apical setae antero-dorsally ( Fig. 33A View Fig ); wing vein R

2+3

with a subapical appendix (e.g., Figs 15F View Fig , 23B View Fig , 25D View Fig ) ............................................ 15

11. Metatrochanter with dark brown transverse carina (as in Fig. 72A View Fig ) .............................................. 12 – Metatrochanter simple, without dark brown transverse carina ( Figs 52B View Fig , 72E View Fig ) ............................ 14

12. Sternum IV posteriorly with 8 very long and strong setae on left side, grouped in two separate groups, the most lateral 3 setae clearly stronger than the 5 more medial ones ( Fig. 42B View Fig ); genitalia, Fig. 42 View Fig C–F ................................................................................................ S. (A.) hansoni Thompson

– Sternum IV posteriorly with 2 or 3 long and strong setae on left side ( Fig. 71B View Fig ) .......................... 13

13. Sternum IV posteriorly on the left side with 2 very long and strong and several very small black setae ................................................................... S. (A.) carinata Hippa, van Steenis & Mutin, 2015

– Sternum IV posteriorly on the left side with 3 rather long and strong and several small black setae ( Fig. 71B View Fig ); genitalia, Fig. 71 View Fig C–E .................................................................... S. (A.) ornata sp. nov.

14. Sternum IV with strong posterior setae on left side only slightly longer than those on right side; sternum VII with a long finger-like process anteriorly and a wide conical tubercle posteriorly ............................................................... S. (A.) index Hippa, van Steenis & Mutin, 2015

– Sternum IV with strong posterior setae on left side much longer and stronger than on right side ( Fig. 16A View Fig ); sternum VII with a large conical and setose tubercle medially ( Fig. 16B View Fig ); genitalia, Fig. 16 View Fig C–E ....................................................................................... S. (A.) bispinosa Brunetti, 1915

15. Wing with crossvein between R 1 and R 2+3 ( Fig. 41F View Fig ); scutellum subtriangular with marginal setae short, about as long as length of scutellum; genitalia, Fig. 44 View Fig B–D ............... S. (A.) hauseri sp. nov.

– Wing without crossvein between R 1 and R 2+3 ( Figs 15F View Fig , 23B View Fig , 25D View Fig ); scutellum subtriangular to slightly semicircular, with marginal setae long, almost twice as long as scutellum ( Fig. 13C View Fig ) ..... 16

16. Metafemur black with basal ¼– ⅓ yellow; scutellum subtriangular, light to dark brown .................. ....................................................................... S. (A.) crassispina Hippa, van Steenis & Mutin, 2015

– Metafemur black with basal ½ yellow; scutellum semi-circular, black; genitalia, Fig. 20 View Fig C–E ......... ...................................................................................................................... S. (A.) clavigera sp. nov.

17. Metatrochanter with dark brown transverse carina ( Fig. 72A View Fig ); sternum VII with a finger-like process anteriorly and a conical tubercle more posteriorly ............................................................................. ............................................................................ S. (A.) malaisei Hippa, van Steenis & Mutin, 2015

– Metatrochanter simple, without carina (e.g., Figs 33 View Fig F–G, 95B); sternum VII without modifications (e.g., Fig. 40A View Fig ), with conical tubercle anteriorly (e.g., Figs 17B View Fig , 37A View Fig , 80A View Fig ) or if with long fingerlike process then without additional conical tubercle (e.g., Figs 27B View Fig , 85A View Fig ) .................................. 18

18. Metafemur with 1–3 long sub-apical setae on antero-dorsal surface ( Fig. 33G View Fig ) ........................... 19 – Metafemur without long sub-apical setae on antero-dorsal surface ............................................... 21

19. Sternum IV posteriorly with 4 evenly long black setae on left side ( Fig. 37B View Fig ); tubercle on sternum VII with normal pile and seta-like pile ( Fig. 37A View Fig ); genitalia, Fig. 37 View Fig C–E ......................................... ...................................................................................................................... S. (A.) gigantea sp. nov.

– Sternum IV posteriorly with 5 long setae on left side, gradually diminishing in length towards medial line or with 3 very long black setae; tubercle on tergum VII with normal pile only ...................... 20

20. Pro- and mesofemur yellow, metafemur yellow on basal ¼– ⅓; vein dm-cu strongly oblique and clearly narrower than other nearby veins; tergum I with 3 black setae at lateral margin; tergum III with anterior ¼ yellow; sternum IV posteriorly on left side with 5 long setae, gradually diminishing in length towards medial line .................................. S. (A.) gigas Hippa, van Steenis & Mutin, 2015

– Pro- and mesofemur black, metafemur black except narrowly yellow at base; vein dm-cu slightly oblique, as wide as other nearby veins; tergum I with 2 black setae at lateral margin; tergum III with anterior ⅓ yellow; sternum IV posteriorly on the left side with 3 very long black setae and some very short ones more medially .................................... S. (A.) trispina Hippa, van Steenis & Mutin, 2015

21. Sternum IV posteriorly on the left side with 2–4 longer setae set apart from other setae and setulae (e.g., Figs 17A View Fig , 27A View Fig , 80B View Fig ) .............................................................................................................. 22

– Sternum IV posteriorly with variable shorter and stronger setulae, without very long setae to the left only ( Figs 40A View Fig , 57 View Fig A–C, 58A) ........................................................................................................ 29

22. Sternum IV posteriorly on the left side with about 4 strong black setae, only slightly longer than other setae on left side, the second seta from left is clearly longer and stronger than the fine ones ( Figs 80B View Fig , 81B View Fig ); sternum VII with long setose rounded tubercle ( Figs 80A View Fig , 81A View Fig ); genitalia, Figs 80 View Fig C–E, 81C– E ........................................................................................................................ S. (A.) setosa sp. nov.

– Sternum IV posteriorly on the left side with 2 or 3 long setae, clearly longer than the fine setae (e.g., Figs 17A View Fig , 27A View Fig , 85B View Fig ); sternum VII with conical tubercle without long setae (e.g., Figs 17B View Fig , 96A View Fig ) or with elongate and apically bent finger-like tubercle ( Figs 27B View Fig , 85A View Fig ) ............................................ 23

23. Sternum IV posteriorly on the left side with 2 longer setae set apart from other setae and setulae (e.g., Figs 17A View Fig , 85B View Fig ); metafemur black and yellow biannulate ..................................................... 24

– Sternum IV posteriorly on the left side with 3 longer setae set apart from other setae and setulae ( Figs 27A View Fig , 96B View Fig ); metafemur bicoloured or biannulate ................................................................... 27

24. Frons with rectangular light grey pollinose vitta along eye-margin ( Fig. 88B View Fig ); sternum IV posteriorly on the left side with the 2 lateral most strong setae twice as long as the more medial ones on the left side ( Fig. 85B View Fig ); sternum VII short pilose, with elongate finger-like process with curved apex ( Fig. 85A View Fig ); ventral lobe of right surstylus simple ( Fig. 85 View Fig C–E) .................. S. (A.) spenceri sp. nov.

– Frons with subtriangular light grey pollinose macula at eye-margin ( Figs 21G View Fig , 88L View Fig ); sternum IV posteriorly on the left side with the 2 lateral most strong setae very long, more than 5 times as long as the other setae ( Figs 17A View Fig , 34B View Fig , 98A View Fig ); sternum VII with a large rounded conical process with a sharp apical projection ( Figs 17B View Fig , 34A View Fig , 98B View Fig ); ventral lobe of right surstylus bifid ( Figs 17 View Fig C–E, 34C–E, 98C–G) ........................................................................................................................................... 25

25. Genitalia, Fig. 98 View Fig C–G: superior lobe on the right side with the longer one of the postero-ventral sublobes long, as long as the main part of superior lobe ...................... S. (A.) vietnamensis sp. nov.

– Genitalia, Figs 17 View Fig C–E, 34C–E: superior lobe on right side with the longer one of the posteroventral sublobes short, much shorter than the main part of superior lobe .................................................. 26

26. Genitalia, Fig. 17 View Fig C–E: superior lobe on the left side with the more anterior one of the postero-ventral sublobes large, several times as long as basally wide, curved, the more posterior one of the sublobes scarcely observable ......................................................................................... S. (A.) bracon sp. nov.

– Genitalia, Fig. 34 View Fig C–E: superior lobe on the left side with the more anterior one of the postero-ventral sublobes small, ca twice as long as basally wide, slightly angled, the more posterior one of the sublobes large ............................................................................................. S. (A.) furcillata sp. nov.

27. Metafemur black and yellow biannulate; postpronotum yellow; genitalia, Fig. 27 View Fig C–E .................... ....................................................................................................................... S. (A.) distincta sp. nov. – Metafemur yellow on basal ¼–1/6 and black apically; postpronotum black ................................... 28

28. Surstyli almost symmetrical ( Fig. 96C, E View Fig ); mesofemur yellow; tergum III with anterior light yellow fascia ( Fig. 97D View Fig ); sternum IV with the longest strong setae longer than the width of sternum ( Fig. 96B View Fig ); genitalia, Fig. 96 View Fig C–E .............................................................. S. (A.) verrucosa sp. nov.

– Surstyli asymmetrical; mesofemur dark brown; tergum III with sub-anterior light yellow fascia; sternum IV with the longest strong setae less than half of the width of sternum ............................................................. S. (A.) trichaeta Hippa, van Steenis & Mutin, 2015

29. Frons black, shiny except for a small elongate densely grey-pollinose macula anterolaterally ( Fig. 54H View Fig ); postpronotum yellow; genitalia, Fig. 58 View Fig B–D ............................... S. (A.) lucida sp. nov.

– Frons black, mainly slightly grey-pollinose with subtriangular densely grey-pollinose macula anterolaterally ( Fig. 38I View Fig ); postpronotum black; genitalia, Figs 40 View Fig B–D, 57D–G ............................ 30

30. Protarsus entirely black; sternum IV with posterior marginal area almost symmetrical, with symmetrical patches of long strong brown setae at posterolateral corner, the setae pale ( Fig. 40A View Fig ); genitalia, Fig. 40 View Fig B–D .................................................................................. S. (A.) hamulata sp. nov.

– Protarsus yellow, with tarsomeres 4 and 5 dark brown to black; sternum IV with posterior marginal area asymmetrical, with asymmetrically placed strong setae or patches of strong setae, the setae at least partly black (e.g., Fig. 57 View Fig A–C) ............................................................................................... 31

31. Scutellar marginal setae rather narrowly separated, distance between setae: length of scutellum 1: 1.4–2.0; genitalia, Fig. 57 View Fig D–G ..................................................................... S. (A.) lobulata sp. nov.

– Scutellar marginal setae very widely separated, distance between setae: length of scutellum 1: 0.9– 1.1 ................................................................................................................................................... 32

32. Gena brown; metafemur with black setulae on yellow basal part; mesofemur dark brown to brown .................................................................... S. (A.) radula Hippa, van Steenis & Mutin, 2015

– Gena yellow; metafemur with or without black setulae on yellow basal part; mesofemur brownyellow to yellow ............................................................................................................................. 33

33. Mesofemur entirely yellow; vein dm-cu slightly oblique to vein M, as wide as other nearby veins, vein CuA 1 extending beyond dm-cu; metafemur with black setulae in yellow basal part .................................................................. S. (A.) raduloides Hippa, van Steenis & Mutin, 2015

– Mesofemur dark brown; vein dm-cu strongly oblique and clearly narrower than other nearby veins, vein CuA 1 not extending beyond dm-cu; metafemur without black setulae in yellow basal part .................................................................. S. (A.) subradula Hippa, van Steenis & Mutin, 2015

34. Metafemur black on apical ½ or more, yellow on basal part, or nearly entirely black; metatibia with large rounded apicoventral dens (e.g., Figs 19F View Fig , 33E View Fig , 52C View Fig ); scutum entirely but weakly greypollinose .......................................................................................................................................... 35

– Metafemur black and yellow biannulate, or femur predominantly yellow; metatibia without apicoventral dens (e.g., Figs 19D View Fig , 33B View Fig ), if with large rounded dens ( Fig. 52F View Fig ) then scutum with clear pollinose pattern ..................................................................................................................... 43

35. Scutellum semi-circular; sternum IV posteriorly on left side or medially with small lobe ( Figs 36A View Fig , 78C View Fig ) ................................................................................................................................................ 36

– Scutellum sub-rectangular; sternum IV posteriorly on left side with large lobe (e.g., Figs 8A View Fig , 14A View Fig , 22B View Fig , 49A) ....................................................................................................................................... 37

36. Vertex narrow, width of head about 6 times as wide as vertex ( Fig. 38F View Fig ); scutellum entirely grey pollinose; postpronotum yellow; sternum IV with weak postero-medial lobe ( Fig. 36A View Fig ); genitalia, Fig. 36 View Fig B–D ....................................................................................................... S. (A.) ghatsi sp. nov.

– Vertex wide, head 4.5 times as wide as vertex ( Fig. 84I View Fig ); scutellum non-pollinose and shiny medially; postpronotum black; sternum IV posteriorly with short, rounded lobe on the left side ( Fig. 78C View Fig ); genitalia, Fig. 78 View Fig A–B, D–F ............................................................................. S. (A.) prolixa sp. nov.

37. Sternum IV posteriorly on the left side with a wide bifid lobe ( Figs 14A View Fig , 22B View Fig 49A) .................... 38 – Sternum IV posteriorly on the left side with a narrow simple lobe (e.g., Figs 8A View Fig , 48B View Fig ) ............... 40

38. Vein dm-cu perpendicular to vein M; the bifid lobe posteriorly on sternum IV with both sublobes approximately equal in size ( Fig. 14A View Fig ); genitalia, Fig. 14 View Fig B–D ........................ S. (A.) bifida sp. nov.

– Vein dm-cu oblique to vein M; the bifid lobe posteriorly on sternum IV with the left side sublobe much narrower than the right side one ( Figs 22B View Fig , 49A) ................................................................ 39

39. The sublobes of the bifid lobe posteriorly on sternum IV separated by a deep cleft (Fig. 49A); genitalia, Fig. 49B–D: dorsal and ventral lobe of surstylus posteriorly weakly separated ..................................................................................................... S. (A.) inventum sp. nov.

– The sublobes of the bifid lobe posteriorly on sternum IV separated by a shallow notch ( Fig. 22B View Fig ); genitalia, Fig. 22 View Fig C–F: dorsal and ventral lobe of surstylus posteriorly separated by a deep notch ............................................................................................................. S. (A.) collicola sp. nov.

40. Postalar callus white to yellow; scutellum sub-rectangular; lobe posteriorly on sternum IV anteriorly narrow, rather straight and directed posteriorly ( Fig. 83A View Fig ); surstyli symmetrical, left surstylus not inflated, genitalia Fig. 83 View Fig B–D ................................................................... S. (A.) spathigera sp. nov.

– Postalar callus black; scutellum semi-circular; lobe posteriorly on sternum IV anteriorly wide and curved laterally ( Figs 8A View Fig , 48 View Fig A–B, 74A); surstyli asymmetrical, left surstylus strongly inflated, genitalia Figs 8 View Fig B–D, 48C–D, 74B–F .............................................................................................. 41

41. Postpronotum black; wing hyaline; genitalia, Fig. 8 View Fig B–D ........................ S. (A.) amplistylus sp. nov. – Postpronotum yellow; wing with weak infuscate pattern; genitalia Figs 48 View Fig C–D, 74B–F .............. 42

42. Wing infuscated at junction of veins R 2+3 and R 4+5 and along vein M 1; genitalia, Fig. 74 View Fig B– F .................................................................................................. S. (A.) philippina Thompson, 1999 – Wing only infuscated at junction of veins R 2+3 and R 4+5; genitalia, Fig. 48 View Fig C–D ................................ .......................................................................................................................... S. (A.) inflata sp. nov.

43. Wing brownish, without darker infuscated pattern; vein dm-cu meeting vein M obliquely ............... ............................................................................... S. (A.) falcata Hippa, van Steenis & Mutin, 2015

– Wing hyaline, infuscated apically and at cross-veins; vein dm-cu meeting vein M perpendicularly ......................................................................................................................................................... 44

44. Wing vein R 2+3 with more or less developed subapical appendix (e.g., Figs 6C View Fig , 23E View Fig , 35B View Fig ) .......... 45 – Wing vein R

2+3

without subapical appendix (e.g., Figs 15B View Fig , 41C View Fig , 45A View Fig ) ........................................ 50

45. Wing without extra crossvein between veins R 1 and R 2+3, at most with a short appendix from vein R 2+3 into cell r 1 ( Fig. 35 View Fig A–B); metafemur evenly curved and with many black setulae on basal yellow part ( Fig. 33D View Fig ) ................................................... S. (A.) forficata Hippa, van Steenis & Mutin, 2015

– Wing with extra crossvein between R 1 and R 2+3 (e.g., Figs 6C View Fig , 23E View Fig ); metafemur straight, without black setulae on basal yellow part (e.g., Fig. 33B View Fig ) ......................................................................... 46

46. Sternum IV posteriorly on the left side with a small subtriangular lobe ( Figs 7A View Fig , 55A View Fig ), if very small and indistinct then several black tooth-like setulae present on posterior part of sternum ( Fig. 87A View Fig ); genitalia: surstyli with clearly differentiated ventral lobe ( Figs 7B, D View Fig , 55B, D View Fig , 87B, D View Fig ) .............. 47

– Sternum IV posteriorly concave, without lobe, without black setulae on posterior part; genitalia: surstyli with hardly differentiated ventral lobe ............................................................................... 49

47. Face and mouth edge black; scutellum medially non-pollinose and shiny; sternum IV with yellowish pile and several black setulae on posterior part ( Fig. 87A View Fig ); genitalia, Fig. 87 View Fig B–D ........................... ..................................................................................................................... S. (A.) strigillata sp. nov.

– Face black, ventral half and mouth edge yellow to dark yellow; scutellum entirely grey pollinose but medially sub-shiny; sternum IV only with light yellow to brown pile ........................................... 48

48. Eye with an area of enlarged facets at anterior margin; pro- and mesotarsus with tarsomeres 2–5 dark brown to black; mesofemur and tibia partly dark brown to black; sternum IV posteriorly with a subtriangular lobe on the left side ( Fig. 7A View Fig ); genitalia, Fig. 7 View Fig B–D .......... S. (A.) albolobata sp. nov.

– Eye without enlarged facets at anterior margin; pro- and mesotarsus with tarsomeres 3–5 dark brown to black; mesofemur and tibia entirely yellow; sternum IV posteriorly with a subtriangular lobe medially ( Fig. 55A View Fig ); genitalia, Fig. 55 View Fig B–D ...................................................... S. (A.) licina sp. nov.

49. Metafemur enlarged, slightly curved basally ( Fig. 33B View Fig ); tergum I with 3 strong setae at lateral margin ( Fig. 13E View Fig ) ......................................................... S. (A.) crucivena Hippa, van Steenis & Mutin, 2015

– Metafemur only slightly enlarged, club shaped; tergum I with 1–2 strong setae at lateral margin ( Fig. 77A View Fig ) ........................................................... S. (A.) mirifica Hippa, van Steenis & Mutin, 2015

50. Basoflagellomere elongate, 1.5 times as long as high; frontal prominence protruding (e.g., Figs 38H View Fig , 54E View Fig , 84C View Fig ); postpronotum yellow to light-brown ............................................................................ 51

– Basoflagellomere short, 1.1–1.3 times as long as high; frontal prominence at most slightly protruding; postpronotum dark brown to black ................................................................................................. 58

51. Scutum with very obvious pollinose pattern with more and less heavy pollinosity; metafemur strongly incrassate ( Fig. 52F View Fig ); crossvein dm-cu meeting vein M slightly obliquely; sternum IV posteriorly on the left side with a short and wide spoon-shaped lobe ( Fig. 53 View Fig A–B); genitalia, Fig. 53 View Fig C–F ................................................................................................. S. (A.) karnataka sp. nov.

– Scutum nearly entirely and evenly pollinose; metafemur slightly incrassate (e.g., Fig. 41C View Fig ); crossvein dm-cu meeting vein M perpendicularly; sternum IV simple (e.g., Figs 12A View Fig , 66A View Fig ) ....................... 52

52. Protarsus not enlarged ( Fig. 61E View Fig ) ................................................................................................... 53 – Protarsus enlarged, Fig. 61 View Fig A–D ...................................................................................................... 54

53. Frontal prominence short ( Fig. 84L View Fig ); protarsus with only tarsomeres 4 and 5 black; wing only infuscated along veins; sternum IV with almost straight posterior margin, without postero-lateral patch of long strong pile ( Fig. 89A View Fig ); genitalia, Fig. 89 View Fig B–D ................... S. (A.) taiwanensis sp. nov.

– Frontal prominence long ( Fig. 84C View Fig ); protarsus with apex of tarsomeres 2 and 3 and entire tarsomeres 4 and 5 black; wing infuscated along veins and an apical macula; sternum IV with posterior margin deeply medially notched, with posterolateral patch of long strong pile ( Fig. 66A View Fig ); genitalia, Figs 66 View Fig B–D, 68B–E, 69A–F ............................................................. S. (A.) orientalis Kertész, 1914

54. Cerci elongate; surstylus large, as long as tergum IX, angle between dorsal and ventral lobe almost right; superior lobe elongate, several times as long as anteriorly wide ( Figs 12 View Fig B–D, 46B–D) ..... 55

– Cerci short oval to trapezoidal; surstylus relatively small, shorter than tergum IX, angle between dorsal and ventral lobe acute; superior lobe subtriangular, at most twice as long as anteriorly wide ( Figs 39 View Fig B–E, 62B–D) ..................................................................................................................... 57

55. Protarsus with tarsomere 3 only medially and tarsomeres 4–5 entirely black; protarsus with basitarsomere short, clearly shorter than combined length of tarsomeres 2 and 3; dorsal lobe of surstylus elongate triangular, narrowing gradually towards apex ...................................................... ............................................................................... S. (A.) nasuta Hippa, van Steenis & Mutin, 2015

– Protarsus with tarsomeres 4 and 5 black and either with tarsomere 3 entirely yellow or entirely black; protarsus with basitarsomere longer, about as long as combined length of tarsomeres 2 and 3; dorsal lobe of surstylus medially constricted, subapically widened ............................................... 56

56. Protarsus with tarsomere 1 yellow, on apical half black and tarsomeres 2–5 entirely black; tarsomere 2 relatively wide, 1.6 times as wide as tibia and only slightly longer than tarsomere 3 ( Fig. 61A View Fig ); genitalia, Fig. 12 View Fig B–D ................................................................................. S. (A.) atrimanus sp. nov.

– Protarsus with tarsomeres 1 and 2 yellow, tarsomere 3 yellow, on apical half black and tarsomeres 4 and 5 entirely black; tarsomere 2 relatively narrow, 1.3 times wider than tibia and almost 1.5 times as long as tarsomere 3 ( Fig. 61C View Fig ); genitalia, Fig. 46 View Fig B–D ..................... S. (A.) incretonigra sp. nov.

57. Protarsus with tarsomere 1 yellow, black on apical half and tarsomeres 2–5 entirely black; tarsomere 1 relatively short, about as long as combined length of tarsomeres 2 and 3; tarsomere 2 very wide, 1.8 times as wide as tibia ( Fig. 61B View Fig ); metafemur anteroventrally with long white pile, almost twice as long as black setae; metatibia without apicoventral dens; only tergum III with anterior yellow fascia; genitalia, Fig. 39 View Fig B–D .................................................................. S. (A.) granditarsis sp. nov.

– Protarsus with tarsomere 1 yellow with black sub-apical annulus, tarsomere 2 dark brown to black with basal ⅓ yellow and tarsomeres 3–5 dark brown; tarsomere 1 relatively long, about as long as combined length of tarsomeres 3, 4 and 5; tarsomere 2 rather wide, 1.3–1.5 times as wide as tibia ( Fig. 61D View Fig ); metafemur anteroventrally with short white pile, only slightly longer than black setae; metatibia with narrow apicoventral dens; terga III and IV with yellow anterior fascia ( Figs 59F View Fig , 63D View Fig ); genitalia, Fig. 62 View Fig B–D .................................................................. S. (A.) nigrotarsata sp. nov.

58. Tergum II dark with anterior yellow fascia; tergum III nearly entirely yellow; thorax light yellow (brownish yellow) with medial and sub-lateral dark brown vittae .... S. (A.) tristriata Brunetti, 1913

– Tergum II black or black with brown to yellow anterior fascia; tergum III black with at most anterior ⅓ yellow; thorax unicolorous dark ................................................................................................. 59

59. Frontal prominence large, clearly protruding beyond eye in profile; protarsus yellow, in some specimens tarsomere 5 brownish; scutellum sub-rectangular ............................................................ ............................................................................... S. (A.) bidens Hippa, van Steenis & Mutin, 2015

– Frontal prominence small, at most slightly protruding beyond eye in profile; protarsus yellow with at least tarsomere 5 black; scutellum semi-circular ........................................................................ 60

60. Face with ventral half yellow to brown-yellow, moderately projected antero-ventrally; protarsus dark brown to black with tarsomeres 1 and 2 yellow; tergum III with anterolateral red macula ( Fig. 90A View Fig ) .......................................................... S. (A.) simplex Hippa, van Steenis & Mutin, 2015

– Face all black, mouth edge sometimes dark brown, very weakly projected antero-ventrally; protarsus yellow with either tarsomeres 4 and 5 or only tarsomere 5 dark brown to black; tergum III with red to yellow anterior fascia ( Fig 70F View Fig ) ................................................................................................. 61

61. Protarsus yellow with tarsomere 5 black, tarsomeres 2–5 slightly widened ( Fig. 61F View Fig ); genitalia, Fig. 76 View Fig B–C ..................................................................................................... S. (A.) plautus sp. nov.

– Protarsus yellow with tarsomeres 4 and 5 dark brown to black, tarsomeres of normal width; genitalia, Fig. 18 View Fig B–C .................................................................................................. S. (A.) brevipilis sp. nov.

62. Metafemur black and yellow biannulate (e.g., as in Figs 6F View Fig , 28B View Fig ) or femur predominantly red to yellow (e.g., Figs 11E View Fig , 95E View Fig ) ........................................................................................................... 63

– Metafemur basally yellow, on apical ½ or more black (e.g., Figs 19F View Fig , 28A View Fig ), in some cases nearly entirely black .................................................................................................................................. 78

63. Wing with infuscate pattern at crossveins and on apex; thorax light yellow with three black vittae on the scutum; sternum IV with a pair of symmetrical sub medial lobes at posterior margin, the lobes with short black spinose setae ( Fig. 11E View Fig ) ........... S. (A.) bilobata Hippa, van Steenis & Mutin, 2015

– Wing hyaline, without infuscate pattern; thorax dark, scutum unicolorous; sternum IV posteriorly unmodified or with one lobe asymmetrically on the left or right side, this lobe without spinose setae ( Figs 9A View Fig , 31A View Fig , 64A View Fig , 94A View Fig ) .............................................................................................................. 64

64. Vein dm-cu meeting vein M perpendicularly; frontal prominence long ( Fig. 38D View Fig ), with distinct medial furrow; basoflagellomere elongate, 2.5 times as long as wide ............................................... .............................................................................. S. (A.) forceps Hippa, van Steenis & Mutin, 2015

– Vein dm-cu meeting vein M obliquely; frontal prominence short ( Figs 30K View Fig , 38A View Fig ) with weak medial furrow; basoflagellomere round to slightly elongate, 1.1–1.5 times as long as wide ..................... 65

65. Sternum IV with large bifid lobe posteriorly on right side ............ S. (A.) amamiensis Shiraki, 1968 – Sternum IV posteriorly without lobe or lobe is placed on the left side .......................................... 66

66. Larger flies, length of body more than 6 mm, pleuron extensively yellow or scutum with clear pollinose pattern ( Figs 15E View Fig , 97B View Fig ); metafemur predominantly orange-yellow ( Fig. 95E View Fig ); terga brownish ......................................................................................................................................... 67

– Smaller flies, length of body less than 5.5 mm, pleuron and scutum uniformly dark brown to black, sometimes scutum with pollinose pattern (e.g., Figs 6 View Fig E–F, 28E, 63A); metafemur dark brown and yellow biannulate; terga black ........................................................................................................ 68

67. Pleuron entirely grey pollinose, on bright orange parts sometimes sub-shiny; arista entirely pilose; sterna III and IV more quadratic, ratio length: width 1: 1.4 respectively 1: 0.60 ....................................................................... S. (A.) cerina Hippa, van Steenis & Mutin, 2015

– Pleuron with at least posterior margin of katepisternum non-pollinose and shiny ( Fig. 65D View Fig ); arista only basally pilose; sterna III and IV more elongate, ratio length: width 1: 2.5–2.7 respectively 1: 0.9–1.1; genitalia, Fig. 94 View Fig B–C ....................................................... S. (A.) varidissima Shiraki, 1930

68. Scutum with pollinose pattern (best viewed dorso-laterally) depending on different density of pollinosity ( Fig. 28E View Fig ); pale markings on tergum III clearly separated from anterior margin ( Figs 10B View Fig , 70B View Fig ) ................................................................................................................................................ 69

– Scutum with uniform dense pollinosity ( Fig. 79F View Fig ); pale markings on tergum III at most very narrowly separated from anterior margin ( Fig. 79F View Fig ) ...................................................................................... 72

69. Pro- and mesoleg entirely yellow .................................................. S. (A.) asciiformis Brunetti, 1915 – Pro- and mesoleg yellow with at least tarsomeres 4 and 5 dark brown to black ............................ 70

70. Scutellar marginal setae almost twice as long as length of scutellum and slightly wider set, 0.13– 0.19 mm apart; protarsus yellow with tarsomeres 3–5 darkened (dark brown to black); genitalia, Fig. 31 View Fig B–C .................................................................................................... S. (A.) farinosa sp. nov.

– Scutellar setae short, at most as long as scutellum and closer set, 0.06–0.08 mm apart; protarsus yellow with tarsomeres 4 and 5 darkened (light-brown to black); genitalia ( Figs 9 View Fig B–E or 64B– D) .................................................................................................................................................... 71

71. Sternum IV simple, with only a weak rounded lobe posteriorly on left side ( Fig. 64A View Fig ); scutum rather uniformly pollinose; genitalia ( Fig. 64 View Fig B–D) ............................................... S. (A.) nubicola sp. nov.

– Sternum IV posteriorly on the left side with large subtriangular lobe ( Fig. 9A View Fig ); scutum pollinose with more densely pollinose submedial vitta; genitalia ( Fig. 9 View Fig B–E) .................................................. ............................................................................................................... S. (A.) apicalis Shiraki, 1930

72. Sternum IV posteriorly on the left side with yellowish sword-like lobe ( Figs 79F View Fig , 82A View Fig ) .............. 73 – Sternum IV posteriorly on the left side without sword-like lobe, at most with short subtriangular lobe ( Figs 63A View Fig , 67C View Fig , 82C View Fig ) ...................................................................................................................... 75

73. Cercus slightly enlarged, shorter than surstylus; gena black .............................................................. .......................................................................... S. (A.) cultrigera Hippa, van Steenis & Mutin, 2015 – Cercus strongly enlarged, as long as or longer than surstylus; gena brown ................................... 74

74. Cercus as long as surstylus; tergum III with red anterior fascia, 1.5 times as long as tergum IV ..................................................................... S. (A.) siculifera Hippa, van Steenis & Mutin, 2015

– Cercus longer than surstylus; tergum III with yellow anterior fascia, 1.1 times as long as tergum IV ........................................................................ S. (A.) ensifera Hippa, van Steenis & Mutin, 2015

75. Cerci asymmetric, greatly enlarged, longer than surstyli; tergum III with anterior yellow fascia ................................................................ S. (A.) sinesmila Hippa, van Steenis & Mutin, 2015

– Cerci symmetrical, slightly enlarged, shorter than surstyli; tergum III with sub-anterior yellow or reddish fascia .................................................................................................................................. 76

76. Sternum IV with conspicuous large subtriangular lobe at posterior margin; metatarsus with tarsomere 1 about 2.5 times as long as wide; frons with shiny fascia posteriorly of lunula; tergum III as long as tergum IV ............................................................ S. (A.) parvula Hippa, van Steenis & Mutin, 2015

– Sternum IV with inconspicuous small lobe at posterior margin; metatarsus with tarsomere 1 about 3 times as long as wide; frons without shiny fascia posteriorly of lunula, entirely grey pollinose; tergum III longer than tergum IV .................................................................................................... 77

77. Tergum III 1.2 times as long as tergum IV; right side surstylus without a third lobe between the dorsal and ventral lobes; left side surstylus with a sub-lobe dorsally at the middle of its dorsal lobe ...................................................................... S. (A.) minuta Hippa, van Steenis & Mutin, 2015

– Tergum III 1.4 times as long as tergum IV; right side surstylus with a third lobe between the dorsal and ventral lobes, this lobe with a deep marginal incision; left side surstylus without a sub-lobe dorsally at the middle of its dorsal lobe ................ S. (A.) pusilla Hippa, van Steenis & Mutin, 2015

78. Larger-sized flies (8.3–8.5 mm); pro- and mesolegs unicoloured brown; terga III and IV unicolorous brownish-black ( Fig. 10A View Fig ) .................................... S. (A.) adusta Hippa, van Steenis & Mutin, 2015

– Smaller-sized flies (4.5–5.5 or 6.5–7.9 mm); pro- and mesolegs at least with basal two or three tarsomeres yellow; terga III and IV dark brown to black with anterior part of tergum III yellow, reddish or brown, paler than the rest of tergum .............................................................................. 79

79. Scutellum shiny, at least medially non-pollinose ............................................................................ 80 – Scutellum dull, pollinose ................................................................................................................ 89

80. Scutum uniformly pollinose; arista long pilose (as in Fig. 43C View Fig ) .................................................... 81 – Scutum slightly grey pollinose with more densely grey pollinose submedial vitta; arista long pilose (as in Fig. 43C View Fig ) or only short pilose basally ( Fig. 43A, D View Fig ) ............................................................ 82

81. Ventral half of face and mouth edge yellow; postpronotum black; katepisternum entirely grey pollinose; genitalia, Fig. 73 View Fig B–C ................................................................. S. (A.) perlobata sp. nov.

– Mouth edge black contrasting with yellow ventral half of face; postpronotum yellow; posterior margin of katepisternum widely non-pollinose, shiny ( Fig. 65A View Fig ); genitalia, Fig. 24 View Fig B–D ................. .......................................................................................................................... S. (A.) crinita sp. nov.

82. Ventral half of face black, edge of mouth dark yellow to black; abdomen non pollinose, shiny black ................................................................................................................................................ 83

– Ventral half of face and edge of mouth yellow; abdomen brown-black to black, slightly pollinose and sub-shiny ......................................................................................................................................... 88

83. Arista entirely long pilose (as in Fig. 43C View Fig ); katepisternum entirely grey pollinose ....................... 84 – Arista short pilose basally, otherwise bare ( Fig. 43A, D View Fig ); posterior margin of katepisternum nonpollinose, shiny ( Fig. 65 View Fig B–C) ......................................................................................................... 85

84. The subtriangular lobe at posterior margin of sternum IV with inconspicuous transverse crest near the base; tergum III and IV asymmetric, their posterior margin skewed to the right; metafemur with only extreme base yellow .................................. S. (A.) atricolor Hippa, van Steenis & Mutin, 2015

– The subtriangular lobe at posterior margin of sternum IV with strong lobe-like transverse crest near the base; tergum III and IV almost symmetric; metafemur with basal 1/6 yellow ..................................................................... S. (A.) furva Hippa, van Steenis & Mutin, 2015

85. Pale fascia on tergum III almost connected to the anterior margin of its tergum ( Fig. 93A View Fig ) ......... 86 – Pale fascia on tergum III clearly positioned sub-anteriorly ( Figs 28B View Fig , 59E View Fig , 63C View Fig ) ......................... 87

86. Arista with basal ¼ rather short pilose ( Fig. 43D View Fig ); medial part of frons along eye-margin widely pollinose, dull ( Fig. 88G View Fig ); genitalia ( Fig. 91 View Fig B–E): cerci dentate ....................................................... ....................................................................................................... S. (A.) tricoloripes Brunetti, 1915

– Arista with basal ⅓ very short pilose; medial part of frons non-pollinose, shiny ( Fig. 88H View Fig ); genitalia ( Fig. 92 View Fig B–C): cerci not dentate ................................................................... S. (A.) umbrosa sp. nov.

87. Katepimeron with anterior half non-pollinose and shiny ( Fig. 65C View Fig ); genitalia ( Fig. 60 View Fig B–C): cerci rounded, simple ................................................................................. S. (A.) nigerrima Shiraki, 1930

– Katepimeron entirely grey pollinose ( Fig. 65B View Fig ); genitalia ( Fig. 26 View Fig C–F): cerci squarish and dentate ............................................................................................................ S. (A.) dentata sp. nov.

88. Pro- and mesofemur predominantly brownish, base and apex narrowly yellow; mesotibia yellow with dark brown apicomedial narrow annulus; metafemur more slender, about 5 times as long as wide ( Fig. 33C View Fig ); scutellum with 2 narrowly separated setae at posterior margin; genitalia, Fig. 29 View Fig B– C ..................................................................................................................... S. (A.) exilipes sp. nov.

– Pro- and mesofemur and tibia entirely yellow; metafemur wider, about 4–4.5 times as long as wide ( Fig. 52E View Fig ); scutellum with 2 widely separated setae at posterior margin; genitalia, Fig. 51 View Fig B– C ........................................................................................................ S. (A.) javana de Meijere, 1914

89. Tergum I on postero-lateral part with almost transverse row of 5 light yellow thin long setae which differ only slightly from surrounding vestiture ( Fig. 13A View Fig ); pro- and mesofemur and tibia at least partly dark ........................................................... S. (A.) achaeta Hippa, van Steenis & Mutin, 2015

– Tergum I on postero-lateral part with an oblique row of 4–5 strong yellow setae which differ greatly from surrounding vestiture (like in Figs 13D View Fig , 47A, D View Fig ); pro- and mesofemur and tibia entirely light yellow ............................................................................................................................................. 90

90. Sternum IV simple with concave posterior margin, without any lobe ( Fig. 32A View Fig ); tergum IV black with yellow fascia anteriorly; genitalia, Fig. 32 View Fig B–F ................................... S. (A.) fimbriata sp. nov.

– Sternum IV posteriorly on left side with subtriangular lobe; tergum IV black or with red anterolateral macula ............................................................................................................................................. 91

91. Pro- and mesolegs entirely yellow; metafemur rather strongly incrassate, 3.4 times as long as wide; tergum III with yellow fascia anteriorly ......................................................................................... 92

– Pro- and mesolegs yellow with at least tarsomere 5 dark brown to black; metafemur slightly less incrassate, 3.6–3.8 times as long as wide; tergum III with red fascia or red lateral macula anteriorly ......................................................................................................................................... 93

92. Thorax entirely heavily grey pollinose; tergum III with yellow anterior fascia ................................. ........................................................................... S. (A.) pollinosa Hippa, van Steenis & Mutin, 2015 – Thorax lightly pollinose, sub-shiny, with more heavily pollinose submedial vitta; tergum III with yellow sub-anterior fascia ...................................................................... S. (A.) tenuis Brunetti, 1915

93. Proleg yellow with tarsomeres 3–5 dark brown; tergum III black with reddish anterior fascia; tergum IV entirely black; sternum IV with large and deep membranous notch posteriorly occupying more than half the length of sternum ............................... S. (A.) culex Hippa, van Steenis & Mutin, 2015

– Proleg yellow with tarsomeres 4 and 5 black; terga III and IV black with yellow anterolateral macula; sternum IV with small membranous notch posteriorly delimited in the posterior marginal area only ................................................................. S. (A.) pollex Hippa, van Steenis & Mutin, 2015

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Syrphidae

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