Jurathauma, Zhang & Shih & Petrulevičius & Ren, 2011

Zhang, Junxia, Shih, Chungkun, Petrulevičius, Julián Fernando & Ren, Dong, 2011, A new fossil eomeropid (Insecta, Mecoptera) from the Jiulongshan Formation, Inner Mongolia, China, Zoosystema 33 (4), pp. 443-450 : 445-448

publication ID	https://doi.org/ 10.5252/z2011n4a2
persistent identifier	https://treatment.plazi.org/id/03AD87D9-F25D-7E4C-F8E2-FA63FBA2FE04
treatment provided by	Felipe
scientific name	Jurathauma
status	gen. nov.

Treatment

Genus Jurathauma n. gen.

Notiothauma View in CoL – M’Lachlan 1877: 427-430 (part).

TYPE SPECIES. — Jurathauma simplex n. gen., n. sp.

ETYMOLOGY. — The generic name is a combination of the word Jurassic and thauma, a Greek word,meaning a wonderful thing or a marvel, from the recent genus of Eomeropidae Notiothauma M’Lachlan, 1877 . Gender feminine.

DIAGNOSIS. — Simplified venation compared to other genera of the family: costal area traversed by one main veinlet and forming two rows of cells; RP + MA with five branches; MP with five branches; CuA double and CuP simple; three anal veins.

Jurathauma simplex n. gen., n. sp. ( Figs 1 View FIG ; 2A, B, D View FIG ; 3 View FIG )

TYPE MATERIAL. — Holotype (CNU-MEC-NN2009001) and paratype (CNU-MEC-NN2009002).

TYPE LOCALITY. — China. Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia; Jiulongshan Formation.

ETYMOLOGY. — The specific name is derived from the Latin simplex (meaning “simple”) referring to its simple venation.

MATERIAL EXAMINED FOR COMPARISON. —Two new specimens described herein: Tsuchingothauma shihi Ren & Shih, 2005 (CNU-M-NN200401 and CNU-M-NN200402) from the Middle Jurassic of Jiulongshan Formation, Daohugou locality, and Typhothauma yixianensis Ren & Shih, 2005 (CNU-M-LB200401) from the Late Jurassic to Early Cretaceous of Yixian Formation, Chaomidian locality.

DISTRIBUTION. — Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China; Jiulongshan Formation.

DESCRIPTION

Based mainly on the holotype, with supplemental information from the paratype.

Holotype

A complete male specimen preserved in dorsal view; wings almost symmetrically arranged, with a pair on each side slightly overlapped; body length (excluding antennae) 15 mm; head and antennae not preserved ( Figs 1A View FIG ; 3A View FIG ).

Paratype

A complete female specimen preserved in dorsal view; forewing and hind wing overlapped and partially preserved; body length (excluding antennae) 20 mm; hypognathous head partially preserved;

antennae incomplete with many antennomeres, filiform ( Figs 2A, B View FIG ; 3B, C View FIG ).

Forewing length 16.1 mm, width 6.2mm; apical margin of the forewing somewhat sharp-pointed ( Fig. 1 View FIG A-C). At very base of the costal margin, a cluster of less than 20 heavy setae (up to 1.35 mm long) for an area of about 1.4 mm wide; distally, an area about 2 mm wide with shorter (up to 0.3 mm long) setae (measurements based on left forewing); costal margin continuing pubescent; costal area narrow basally to humeral vein, distally broadened, traversed by two rows of cells and a strong and longitudinal secondary vein parallel to the costal margin with four (in right wing) and seven (in left wing) crossveins; three crossveins from ScP to the secondary longitudinal vein; ScP distally vanishing and not reaching anterior wing margin; pterostigma present, apical part of RA vanishing or obscure in this area; RA, RP + MA and MP forming the “ Eomeropidae triadic branching” ( Fig. 1B View FIG ); RA diverging anteriorly from RP + MA, as in Tsuchingothauma shihi Ren & Shih, 2005 ( Fig. 2D View FIG ); RP + MA continuing straight with R; RA simple, weakly zigzagged; RP + MA with two main branches, anterior one with three terminal branches, posterior one with two terminal branches; three setae on Cu just before its bifurcation, a longer seta in the beginning of CuA; MP bifurcated into two main branches, MP1 + 2 continuing the same direction, MP3 + 4 diverging posteriorly at a nearly straight angle; MP1 single, MP2 double, MP3 single, MP3 + 4 joined with CuA1 + 2 by a short distance in left wing, MP3 + 4 only touching CuA1 + 2 in right wing (see arrows in Figure 1B, C View FIG ); CuA1 + 2 free (?); CuA3 + 4 strongly and curvy zigzagged by crossveins, bent sharply posteriad approximately at branching of MP, only with one terminal branch; CuP single, zigzagged by crossveins; three simple anal veins. All veins thickened in basal parts, then narrower; no coloration detected. Entire forewing with many crossveins and long hexagonal cells.

Hind wing length 12.5 mm, width 5.0 mm; veins partly preserved ( Fig. 1A, D View FIG ). Costal area with few (at least three) crossveins; pterostigma present; apical part of RA vanished or obscure in this area; RP + MA arising from RA at acute angle; RA continuing stem of R; RA simple, posteriorly limiting pterostigma; RP + MA divided in two main branches, anterior one with three terminal branches, posterior one with two terminal branches; MP1 single, MP2 double, MP3 single, MP3 + 4 only touching CuA1 + 2 in right wing (as in recent Notiothauma ); CuA incompletely preserved. All veins thickened in basal parts, then narrower; no trace of coloration. Compared to the forewing, the hind wing smaller in size and with fewer crossveins and long hexagonal cells.

Prothorax poorly preserved. Meso- and metanotum distinctly larger than pronotum. There is a certain amount of crinite on the thorax ( Figs 1A View FIG ; 3A, B View FIG ).

Legs densely clothed with transverse rows of short setae. Femora stout.Tibiae somewhat longer and slender, with many conspicuous spines and 2 terminal spurs. Tarsi 5-segmented, basitarsus longest. Pretarsus with 2 lateral claws ( Figs 2C View FIG ; 3D, E View FIG ).

Abdomen elongate, tapering apically, with 9 visible segments. Segments 8 and 9 abruptly more slender than 1-6. Lateral tergal horns in segment 7. Gonostylus partially preserved ( Fig 1A View FIG ).