Scydmaenilla (Zeanilla) newtoni (Franz) Franz, 2015

Jałoszyński, Paweł, 2015, Revision of the genus Stenichnaphes Franz in New Zealand (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 3915 (2), pp. 250-262 : 259-261

Scydmaenilla (Zeanilla) newtoni (Franz) , comb. n.

( Figs. 3, 6 , 18–23 , 26 )

Stenichnaphes newtoni Franz, 1986: 197 , Fig. 1 .

Type material. Holotype: NEW ZEALAND: ♂, five labels ( Fig. 6 ): " NEW ZEALAND: MB / Pelorus Bridge / Scenic reserve, 60 m / 15.xii.1984 - 4.i.1985 / hdwd.-podocarp forest / A.Newton/M.Thayer 711" [white, printed], "window / trap" [white, printed], " Stenichnaphes / newtoni m. / det.H.Franz" [white, handwritten and printed], " Holotypus " [red, handwritten], and white printed NZAC catalogue number (04070025) and QR code ( NZAC).

Additional material studied (4 exs): 1 ♂, New Zealand, Lynfield, 21.vi.1981, Salix fragilis wood mould, leg. G. Kuschel; 1 ♀, New Zealand, Napier, 22.x.1984, pohutukawa [i.e., Metresideros excelsa ], leg. C.F. Butcher; 1 ♀, New Zealand, Bethells, Matuku Res., 1.v.-28.ix.1988, Malaise trap on tree platform; 1 ♀, New Zealand, Poor Knights Is., Tawhiti Rahi, 7.xii.1980, sifted litter and decayed wood, leg. G. Kuschel (specimens in NZAC and cPJ).

Revised diagnosis. This is the only species of Zeanilla and can be identified on the basis of subgeneric diagnostic characters (first of all a pair of long lateral pronotal furrows) and aedeagus with a complicated symmetrical system of elongate and pointed sclerites projecting from ostium.

Redescription. Body of male ( Fig. 3 ) convex, reddish-brown, covered with light brown vestiture; BL 1.25–1.38 mm.

Head ( Figs. 3 , 18 ) broadest at large, strongly convex and moderately coarsely faceted eyes, HL 0.20 mm, HW 0.25 mm; tempora in dorsal view distinct, as long as only 1/3 of length of eye; vertex and frons confluent and weakly convex; supraantennal tubercles feebly marked, anterior margin of frons subtriangular. Frons and vertex appear impunctate under magnification 80×; setae long and sparse, suberect to erect. Antennae slender, AnL 0.58–0.60 mm; antennomeres I–II distinctly elongate, III–IV slightly elongate, V–VI about as long as broad or barely discernibly transverse, VII–X distinctly transverse, XI about 1.5× as long as broad, with rounded apex.

Pronotum ( Figs. 3 , 19 ) distinctly elongate, broadest at base; PL 0.30–0.33 mm, PW 0.28–0.29 mm. Anterior and lateral margins in anterior third rounded; sides slightly concave in posterior third; posterior margin arcuate; lateral longitudinal furrows, antebasal transverse groove and lateral pits distinct. Punctures on pronotal disc as inconspicuous as those on head; setae sparse, long and erect.

Elytra ( Fig. 3 ) distinctly more convex than pronotum, oval, broadest distinctly anterior to middle; EL 0.75–0.85 mm, EW 0.53–0.58 mm, EI 1.43–1.48; humeral calli distinct and elongate, basal impression on each elytron short but distinct; apices separately rounded, elytra slightly impressed along suture near anterior third. Punctures on elytra slightly more distinct than those on pronotum but still barely discernible under magnification 80×; setae sparse, much longer than those on pronotum, erect. Hind wings well developed.

Legs ( Fig. 3 ) long and slender; all tibiae nearly straight.

Aedeagus ( Figs. 22–23 ) stout and thin-walled; AeL 0.23–0.25 mm; median lobe and endophallus symmetrical, apical projections short; endophallic structures complicated and darkly sclerotized, with subapical group of several long and pointed sclerites directed distally and lateral groups of submedian dense and thin sclerites directed distomesally. Parameres slender, each with two apical setae.

Female. Externally indistinguishable from male. BL 1.25–1.26 mm; HL 0.18 mm, HW 0.25 mm, AnL 0.53 mm; PL 0.30–0.31 mm, PW 0.28 mm; EL 0.78 mm, EW 0.55–0.63 mm, EI 1.24–1.41.

Distribution. North Island (including Poor Knights Islands) and the northern tip of South Island, New Zealand ( Fig. 26 ).

Figures

FIGURES 1 – 6. Dorsal habitus (1 – 3) and labels of holotypes. Stenichnaphes urbanus Franz (1, 4); Stenichnaphes peloriensis Franz (2, 5); Scydmaenilla newtoni (Franz) (3, 6).

FIGURES 18 – 23. Morphology of Scydmaenilla newtoni (Franz). Head in ventral view (18), prothorax in dorsal (19) and ventral (20) view, pterothorax in ventral view (21), aedeagus in ventral (22) and lateral (23) view. Abbreviations: abg, antebasal pronotal groove; ar, anterior ridge; bs, basisternal part of prosternum; bst, basistipes; cd, cardo; elp, external lateral pit; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hy, hypomeron; ilp, internal lateral pit; lac, lacinia; lf, lateral pronotal furrow; lmfa, lateral metafurcal arms; lp., labial palp; ls, lateral suture of submentum; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; mst, mediostipes; msvp, mesoventral process; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, pronotosternal suture; occ, occipital constriction; par, procoxal cavity; pcr, procoxal rest; pcs, procoxal socket; pf, profurca; pl, posterior lobe; pmn, prementum; pre, prepectus; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; sp, spermatheca; tm, temple; v 3, metaventrite.

FIGURES 24 – 26. Distribution of Stenichnaphes urbanus Franz (24), Stenichnaphes peloriensis Franz (25) and Scydmaenilla newtoni (Franz) (26).