Anisepyris Kieffer

Waichert, Cecilia & Azevedo, Celso O., 2009, Phylogenetic analysis of Rhabdepyris (Hymenoptera: Bethylidae) and redefinition of generic limits based on morphological characters, Zootaxa 2284, pp. 1-29 : 22-25

Anisepyris Kieffer

Anisepyris Kieffer, 1905 , In André, Spec. Hymen, Eur. Alger., 9: 248. Type species: Epyris amazonicus Westwood. Monotypic designation.

Rhabdepyris (Lophepyris) Evans, 1959 , Proc. Ent. Soc. Wash., 61: 201–204. Type species: Rhabdepyris (Lophepyris) bridwelli Evans. Original designation. Synonym designation by Evans, 1964, Boll. Mus. Comp., 132: 96. [Not examined].

Trichotepyris Kieffer, 1906, In André, Spec. Hymen. Eur. 9: 376. Type-species: Rhabdepyris pallidipennis Kieffer. Subsequent designation by Muesebeck & Walkley 1951, In C. W. F. Muesebeck, K. V. Krombein & H. K. Townes, Hymen. Syn . Cat., 2: 729 (proposed as subgenus of Rhabdepyris ). Synonym designation by Kieffer, 1914, Das Tierreich, 41: 346. Subgenus designation by Evans, 1965, Boll. Mus. Comp., 133: 69. Syn . nov. [Not examined].

Diagnosis. It is usually black or dark metallic in coloration; with large mandible, from 2 to 5 teeth; clypeus with median lobe usually angulated, median carina arched, and lateral lobes not developed; eyes large and densely pilose; antennae arising from lower side of a transverse prominence formed by lower front; malar space margined by a carina. Pronotum usually with a transverse carina in front, disc large flat or weakly convex, its sides rather sharp and often also carinate, pronotal disc often with a foveolate groove in the posterior portion; notauli sinuous, complete or nearly so; scutellum with a transverse basal groove which is fairly wide and commonly turned backward and slightly enlarged on each side; propodeal disc subquadrate, with strong lateral and posterior carinae, the postero-lateral corners often foveolate, median carina complete and extending down the declivity; mesopleuron with prepectal and postpectal carinae, generally, the lower fovea is present, mesopleural fovea elongate, episternal groove is present and scrobiculate.

Comments and character discussion. Kieffer (1905) recognized Anisepyris primarily by the anterior carina of the pronotum (character 20), the transverse groove on the scutellum (24), and the number of flagellomeres of the antenna. Herein, we also include the following characters: head and eyes large and densely pilose (12, 13), the clypeus with an arched median carina (5), the pronotal disc with a foveolate groove in the posterior portion (24), and the notauli generally sinuous, fine and shallow (31). Often, the species of this genus are also shiny (46) and have a malar space that is margined by a carina (6). The body shape is similar to Chlorepyris . Yet Anisepyris can be more carinate and marked with fovea and grooves, and the tarsal claws are distinct; Chlorepyris species do not have a carinate malar space.

Species of Rhabdepyris (Rhabdepyris) transferred to Laelius

Nomenclatural Nomenclatural

Species Species

act act

* areolatus ( Rosmann & Azevedo, 2005) comb. nov. * minutulus (Evans, 1965) comb. nov. * canchinensis (Azevedo, 1992) comb. nov. * muesebecki (Evans, 1965) comb. nov. * gracilis (Evans, 1965) comb. nov. * multilineatus (Evans, 1966) comb. nov. * hirticulus (Evans, 1965) comb. nov. * nigripilosus (Ashmead, 1895) comb. nov. * maboya (Snelling, 1996) comb. nov. * nigriscapus (Evans, 1965) comb. nov. * mellipes (Evans, 1965) comb. nov. * versicolor (Evans, 1970) comb. nov.

Species of Rhabdepyris (Trichotepyris) transferred to Anisepyris Species of Rhabdepyris (Chlorepyris) transferred to Chlorepyris stat. rev.

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Species of Rhabdepyris (Chlorepyris) transferred to Chlorepyris stat. rev.

Mesopleuron ( Figs. 4 C, 4F). It seems to be surrounded by the prepectal (47) and postpectal carina (48). Generally, the postpectal carina is closed and forms the lower fovea (49, 50); the mesopleural fovea is elongate (52) and, some of species has a triangular shape; the mesopleural pit is deep. The episternal groove is present (55) and scrobiculate (56) in some species, the sub-tegular groove is complete; the anterior-superior fovea (53) is rounded and deep, in median to large size (54).

Variation. The dorsal profile of the temples may be long and more curved in some specimens; the anterior transverse carinae on pronotal disc may be absent. Some species have the notauli shorter and dilated in the posterior portion. Mesopleuron may have small fovea replete with punctuation (mainly inferiorly) that often confuse the limits or the delimiting carinae. Specimens usually classified as Trichotepyris do not have anterior carina on pronotum.

Figures

FIGURE 4. A. E. niger: mesopleuron. B. E. blandus: mesopleuron. C. A. amazonicus: mesopleuron. D. Rhabdepyris sp. 4: mesopleuron. E. B. aurata: mesopleuron. F. A. smithanus: mesopleuron. Scale = 0.35 mm.